A remarkable fact in the distribution of European Snakes is the
altitudinal range of _Vipera berus_, _V. aspis_, and _V. ursinii_. The
first being the northernmost snake, generally distributed in Northern
Europe and more locally in the south, should, one would expect, be a
mountain form in the south. This is so in Switzerland, where it occurs
chiefly between 2,500 and 9,000 feet, on the northern aspect of the
Alps, whilst _V. aspis_ lives at altitudes below 5,000 feet; but on the
southern aspect of the same chain things are reversed, and _V. berus_ is
replaced by _V. aspis_, which reaches an altitude of 9,700 feet, whilst
the former shows a tendency to abandon the mountains, and has
established itself in a few localities in the plain of North Italy.
Again, in France _V. berus_ is the northern and _V. aspis_ the southern
species, yet the latter is the only one found on the French side of the
Pyrenees (up to 7,250 feet), whilst the former reappears in
North-Western Spain and Portugal at very low altitudes, even at
sea-level. _V. ursinii_ is a mountain form in Italy (Abruzzi), in France
(Basses-Alpes), and in the Balkan Peninsula (up to 6,800 feet); but it
is restricted to the plain in Lower Austria and Hungary, where _V.
berus_ occurs only in the mountains.
Only three species are entirely confined to Europe:
_Coluber scalaris_, _Vipera ursinii_, and _V. aspis_.
Of the species which range outside Europe, the following occur both in
Western Asia and in North Africa:
_Eryx jaculus_, _Tropidonotus natrix_, _Coelopeltis monspessulana_,
_Vipera lebetina_.
In Western Asia and the North-East of Egypt:
_Tropidonotus tessellatus_, _Zamenis dahlii_.
In Western Asia:
_Typhlops vermicularis_, _Zamenis gemonensis_, _Coluber
quatuorlineatus_, _C. dione_, _C. longissimus_, _C. leopardinus_,
_Coronella austriaca_, _Contia modesta_, _Tarbophis fallax_, _T.
iberus_, _Vipera renardi_, _V. berus_, _V. ammodytes_, _Ancistrodon
halys_.
In North Africa:
_Macroprotodon cucullatus._
In North-West Africa:
_Tropidonotus viperinus_, _Zamenis hippocrepis_, _Coronella girondica_,
_Vipera latastii_.
The following lists will help to elucidate the distribution of the
snakes in the different parts of Europe:
I. SCANDINAVIA
1. _Tropidonotus natrix_ (as far north as 65°).
2. _Coronella austriaca_ (as far north as 63°).
3. _Vipera berus_ (as far north as 67°).
II. GREAT BRITAIN
1. _Tropidonotus natrix_ (England and Wales, extreme south-east of
Scotland).
2. _Coronella austriaca_ (Surrey, Berkshire, Hampshire, and
Dorsetshire).
3. _Vipera berus._
III. BELGIUM AND HOLLAND
1. _Tropidonotus natrix._
2. _Coronella austriaca._
3. _Vipera berus._
IV. GERMANY AND DENMARK
1. _Tropidonotus natrix._
2. _Tropidonotus tessellatus_ (Middle Rhine and Moselle, Saxony).
3. _Coluber longissimus_ (Denmark, Schlangenbad, Treves).
4. _Coronella austriaca._
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