The Social Direction of Evolution: An Outline of the Science of EugenicsKellicott, William E. (William Erskine)
Science
The Social Direction of Evolution: An Outline of the Science of Eugenics
Kellicott, William E. (William Erskine)
Eugenics
Turning now to variation of the second type we find what are
ordinarily called _mutations_, or differences quite properly termed
_variations_, in a strict sense, as distinguished from the preceding
fluctuations or variability phenomena. Mutations or variations are
abrupt changes of the average or type condition to a new condition or
value which then becomes a new center of fluctuating variability. The
difference between variability and variation may be illustrated
through an analogy suggested by Galton (Fig. 5). A polygonal plinth,
or better a polyhedron, resting upon one face is easily tipped
slightly back and forth, but after slight disturbance it always
returns to its first position of stable equilibrium. Each face of the
plinth or polyhedron represents an organismal characteristic; these
slight backward and forward movements represent fluctuations, always
centering about the average condition. An unusually hard push sends
the plinth over upon another face in which it has a new position of
stability; this represents true variation or mutation. In this new
position it is again stable, may again be rocked back and forth
showing fluctuations about its new average position.
[Illustration: FIG. 5.--Plinth to illustrate the difference
between variability (fluctuation) and variation (mutation).]
The essential difference between true variation and fluctuation or
variability of an extreme nature, is with reference to the inheritance
of such divergence. In the second generation the offspring of extreme
variates or fluctuations have not the same average as their own
parents but an average much nearer that of the whole group to which
their parents belonged; the average stature of the children of
unusually short or tall parents is respectively greater or less than
that of their own parents--that is, is nearer the average of the whole
group of parents, provided the shortness or tallness of the parents is
a fluctuation. When the shortness or tallness is a true variation or
mutational character, offspring have approximately the same average
stature as their immediate parents, although the children of course
show fluctuation in height so that some are slightly above and others
slightly below the parental height.
Mutations may occur through the addition or the subtraction of single
characters of the simple or unit type. Such are the variations from
brown or blue eyes to albino, five fingers to six, and the like. These
are the familiar "sports" of the horticulturalist and breeder. They
are of the greatest value in evolution, for it seems quite likely that
it is only through the permanent racial fixation of these mutations
that permanent changes in the characters of a breed may be effected,
i. e., evolution occurs primarily through mutation.
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