The Story of a Boulder; or, Gleanings from the Note-book of a Field GeologistGeikie, Archibald
Science
The Story of a Boulder; or, Gleanings from the Note-book of a Field Geologist
Geikie, Archibald
Geology; Petrology
The body of the megalichthys was cased in an armature of as solid and
glittering bone as that which defended its head. Where the plates of
the cranium ended off they were succeeded by large rhomboidal scales
that crossed the body obliquely, and overlapped each other like the
metal plates in the antique scale-armour. Each scale consisted of
two parts, of which one had a rhomboid form and was covered over with
enamel, while the other ran round the two inner sides of the rhomb
as a broad unenamelled selvage deeply indented along its centre. It
was the enamelled portion alone that formed the outer surface, the
rough unpolished border being covered by the overlapping edges of
the adjoining scales. The scales had not a uniform thickness, but
were strongest at the covered part from which each thinned off to
the outer edges. In this way the thin edge of one scale pressed down
on the thick part of the subjacent one, and a covering of uniform
strength and smoothness was produced. Looking at a set of these scales
as they still occupy their original position on the creature's body,
it is scarcely more than a half of each which meets our eye; for the
unenamelled border occupied about a third of the entire surface, and
a fourth of the remainder was covered by the overlapping scales. The
effect of this arrangement must have been to combine great strength
with the most perfect flexibility. Notwithstanding the bulk of his
helmet and the weight of his scale-armour, we cannot conceive the
megalichthys to have been other than a lithe, active, predaceous fish,
dealing death and destruction among the herring-like shoals of little
palæonisci and amblypteri, though able to maintain perhaps but a
doubtful warfare with his more bulky contemporary, the holoptychius.
The internal structure of the scales of the megalichthys exhibits the
same provision for combining strength with the least possible amount
of material. Viewed in a transverse section under a magnifying power
of about eight diameters, they are seen to consist of three layers of
bone; each possessing a peculiar structure. The outermost is formed
of a tessellated pavement of minute round ocelli, having a fine brown
colour, and placed close together with considerable regularity. They
somewhat resemble little wheels, the axle being either a dark solid
nucleus or a small circular aperture, whence there radiates to the
outer rim a set of exceedingly minute fibres which were originally
hollow, and served as canals to carry on the growth of the scale. The
vacant space left where four wheels impinge on each other, forms one
of the pores that cover the enamelled surface of the scale. The whole
structure of this outer layer very closely resembles that presented by
the internal part of the base of the teeth, save that the confluent
lobes shown in the teeth become in the scale detached into separate and
independent circles. The central stratum of each scale is composed of
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