The Story of a Loaf of BreadWood, T. B. (Thomas Barlow)
Science
The Story of a Loaf of Bread
Wood, T. B. (Thomas Barlow)
Bread; Wheat
In order to avoid these difficulties the method of selection according
to progeny has been devised. The essence of this method is to select
for stock, not the best individual plant, but the plant whose progeny
yields the greatest weight of seed per unit area. This method was
applied with great industry and some success in the Minnesota wheat
breeding experiments of Willett Hays. Large numbers of promising plants
were collected from a plot of the best variety in that district. The
seed from each plant was rubbed out and sown separately. One hundred
seeds from each plant were sown on small separate plots which were
carefully marked out and labelled. Every possible precaution was taken
to make all the little plots uniform in every way. By harvesting each
plot separately, and weighing the grain it produced, it was possible to
find out which of the original plants had given the largest yield. This
process was repeated by sowing again on separate plots a hundred seeds
from each individual plant from the best plot, and again weighing the
produce of each plot. After several repetitions it was stated that new
strains were obtained which yielded considerably greater crops than the
variety from which they were originally selected. These results were
published in 1895, but no definite statements have since appeared as to
the success ultimately attained.
This method of selection is undoubtedly more likely to give successful
results than the method which depends on the selection of plants for
their apparent good qualities; but it has several weak points. In
the first place it is almost impossible to make the soil of a large
number of plots so uniform that variation in yield due to varying soil
conditions will not mask the variations due to the different cropping
power of the seed of the separate plants. Many experimenters are still
at work with a view to overcome this difficulty. Secondly, plant
breeders are by no means agreed on the exact theoretical meaning of
improvement by selection. The balance of evidence at the present time
seems to tend towards the general adoption of what is known as the
pure-line theory. According to this theory, which was first enunciated
by Johannsen of Copenhagen as the outcome of a lengthy series of
experiments with beans, the general population of plants, in say a
field of wheat of one of the standard varieties giving an average
yield of say 40 bushels per acre, consists of a very large number of
races each varying in yielding capacity from say 30 to 50 bushels per
acre. These races can be separated by collecting a very large number
of separate plants, sowing say 100 seeds from each on a separate plot,
and weighing the produce separately. The crop on each plot, being the
produce of a separate plant, will be a distinct race, or pure line
as it is called, and each pure line will possess a definite yielding
power of its own. If this is so the difficulty of soil variation can be
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