Calamites are among the most abundant fossil plants of the Carboniferous period, and occur also in the Devonian. They are preserved as striated, jointed, cylindrical, or compressed stems, with fluted channels or furrows at their sides, and sometimes surrounded by a bituminous coating, the remains of a cortical integument. They were originally hollow, but the cavity is usually filled up with a substance into which they themselves have been converted. If, during the coal-period, the vegetable kingdom had reached its maximum, the animal kingdom, on the contrary, was poorly represented. Some remains have been found, both in America and Germany, consisting of portions of the skeleton and the impressions of the footsteps of a Reptile, which has received the name of Archegosaurus. Among the animals of this period we find a few Fishes, analogous to those of the Devonian formation. These are the _Holoptychius_ and _Megalichthys_, having jawbones armed with enormous teeth. Scales of _Pygopterus_ have been found in the Northumberland Coal-shale at Newsham Colliery, and also in the Staffordshire Coal-shale. Some winged insects would probably join this slender group of living beings. It may then be said with truth that the immense forests and marshy plains, crowded with trees, shrubs, and herbaceous plants, which formed on the innumerable isles of the period a thick and tufted sward, were almost destitute of animals. Coal, as we have said, is only the result of a partial decomposition of the plants which covered the earth during a geological period of immense duration. No one, now, has any doubt that this is its origin. In coal-mines it is not unusual to find fragments of the very plants whose trunks and leaves characterize the Coal-measures, or Carboniferous era. Immense trunks of trees have also been met with in the middle of a seam of coal. In order to explain the presence of coal in the depths of the earth, there are only two possible hypotheses. This vegetable débris may either result from the burying of plants brought from afar and transported by river or maritime currents, forming immense rafts, which may have grounded in different places and been covered subsequently by sedimentary deposits; or the trees may have grown on the spot where they perished, and where they are now found.