The Structure and Life-history of the Cockroach (Periplaneta orientalis): An Introduction to the Study of InsectsMiall, L. C. (Louis Compton)
Science
The Structure and Life-history of the Cockroach (Periplaneta orientalis): An Introduction to the Study of Insects
Miall, L. C. (Louis Compton)
Cockroaches
The lower or less-specialised Insects, such as the Cockroach, have
nearly as many nerve-ganglia as segments, and the longitudinal
connectives of the nerve-cord are double. In the adult of certain
higher Insects[9] (_e.g._, many Coleoptera, and some Diptera) the
nerve-ganglia are concentrated, reduced in number, and restricted to
the head and thorax; while all the connectives, except those of the
œsophageal ring, may be outwardly single.
[9] So also in some larvæ (_Calandra_, _Œstrus_, &c.).
The heart, or dorsal vessel, is subdivided by constrictions into a
series of chambers, from which an aorta passes forwards to the head.
Air is usually taken into the body by stigmata or breathing-pores,[10]
which lie along the sides of the thorax and abdomen. It circulates
through repeatedly-branching tracheal tubes, whose lining is
strengthened by a spiral coil. Air-sacs (dilated portions of the
air-tubes) occur in Insects of powerful flight.
[10] In some aquatic Insects the exchange of gases is effected by
“pseudobranchiæ,” and the tracheal system is closed.
The generative organs are placed near the hinder end of the body.[11]
Most Insects are oviparous.[12] The sexes are always distinct; but
imperfect females (“neuters”) occur in some kinds of social Insects.
Agamogenesis (reproduction by unfertilised eggs) is not uncommon.
[11] Dragon-flies have the male copulatory apparatus, but not the
genital aperture, in the fore part of the abdomen.
[12] Aphis and Cecidomyia are at times viviparous, and a viviparous
Moth has been observed by Fritz Müller (Trans. Entom. Soc. Lond.,
1883).
_Orders of Insects._
The orders of Insects are usually defined with reference to the
degree of metamorphosis and the structure of the parts of the mouth.
Five of the orders (3, 5–8) in the table on page 9 undergo complete
metamorphosis, and during the time of most rapid change the insect
is motionless. In the remaining orders (1, 2, 4) there is either no
metamorphosis (_Thysanura_), or it is incomplete--_i.e._, the insect
is active in all stages of growth. Among these three orders we readily
distinguish the minute and wingless Thysanura. Two orders remain,
in which the adult is commonly provided with wings; of these, the
Orthoptera have biting jaws, the Hemiptera, jaws adapted for piercing
and sucking.
The name of Black Beetle, often given to the Cockroach, is therefore
technically wrong. True Beetles have a resting or chrysalis stage, and
may further be recognised in the adult state by the dense wing-covers,
meeting along a straight line down the middle of the back, and by the
transversely folded wings. Cockroaches have no resting stage, the
wing-covers overlap, and the wings fold up fan-wise.
_Further Definition of Cockroaches._
In the large order of Orthoptera, which includes Earwigs, Praying
Insects, Walking Sticks, Grasshoppers, Locusts, Crickets, White Ants,
Day-flies, and Dragon-flies, the family of Cockroaches is defined as
follows:--
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