The Structure and Life-history of the Cockroach (Periplaneta orientalis): An Introduction to the Study of InsectsMiall, L. C. (Louis Compton)
Science
The Structure and Life-history of the Cockroach (Periplaneta orientalis): An Introduction to the Study of Insects
Miall, L. C. (Louis Compton)
Cockroaches
A certain amount of geological evidence may also be claimed in support
of this view. A survey of the species of the two groups found up to
the present time in America, published and unpublished, shows that all
the _Mylacridæ_ are found below the Upper Carboniferous, while more
than half the _Blattinariæ_ are found in or above it. This results
largely from a recent and as yet unpublished discovery of _Blattinariæ_
in the Upper Coal Measures of Ohio and West Virginia, which in their
general features are much nearer than previously discovered American
Cockroaches to the European _Blattinariæ_, the latter of which come
generally from Upper Carboniferous beds. The _Mylacridæ_ have therefore
been found in America in strata generally regarded as older than those
which in Europe have yielded Cockroaches, and this gives a sufficient
explanation why no _Mylacridæ_ have yet been found in the Old World.
In America one is mostly dealing with absolutely older forms, and they
naturally give that continent a more old-fashioned look, when we regard
the Carboniferous fauna as a whole. As already stated, a wing from the
French Silurian (_Palæoblattina Douvillei_ Brongn.) has been claimed as
a Cockroach, but without good reason, and to see a real old Cockroach
one must look to America.
Up to this point we have contrasted the palæozoic Cockroaches with the
existing forms only, and finding such important distinctions between
them, we naturally turn with some curiosity to the intermediate
mesozoic and tertiary formations.
Now, not only are the mesozoic species as numerous (actually, but not
relatively) as the palæozoic, but a recent discovery of a Triassic
fauna of considerable extent, in the elevated parks of Colorado,
presents us with a series of intermediate forms between those peculiar
to the Coal Measures and those characteristic of the later mesozoic
rocks. Excluding, however, for a moment this Triassic fauna, we may
say of the later mesozoic species that they are _Neoblattariæ_, not
_Palæoblattariæ_, though they still show some lingering characteristics
of their ancestry. Thus the front wings are in general of a less
dense texture than in modern times, but without the perfect
diaphaneity of the palæozoic species; in some the anal veins fall in
true palæoblattarian fashion on the inner margin, while in others
which cannot be dissociated generically from them, the anal veins
are disposed as in modern types. But in all there is a loss of one
of the principal veins, or rather an amalgamation of two or more--a
characteristic of more fundamental character. As a general rule,
moreover, to which we shall again advert, the mass of the species are
of small size, in very striking contrast to the older types.
[Illustration: Fig. 122.--_Neorthroblattina Lakesii_ Scudd. × 5. Trias,
Colorado.]
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