The Subspecies of the Mountain ChickadeeGrinnell, Joseph
Science
The Subspecies of the Mountain Chickadee
Grinnell, Joseph
Birds -- California; Chickadees
Close scrutiny of the series of specimens at hand well representing the
entire Rocky Mountain area reveals no variation in phylogenetic
characters from the northernmost to the southernmost stations. All show
in apparently equal degree the long tail and cinnamon tinge of sides and
back, these features together constituting the grounds for separate
subspecific recognition of a Rocky Mountain form. On the other hand, the
Sierra Nevadan center, with its own recognizable race, [Page 507] of
relatively short tail, proves to have two outlying divergent forms.
These three forms are alike in their lack of any cinnamon tinge, this
being replaced in two of them by a buffy tinge and in one form by leaden
gray. The tail in one of the outlying forms is long, in the other short.
The habitats concerned are, respectively, the desert mountains of the
Inyo region of eastern California, and the coastal mountains of southern
California. This differentiation within the Pacific district,
particularly within the state of California, will be better understood
in its geographic bearing by reference to the accompanying map (fig. 1).
The behavior of the tail of _Penthestes gambeli_--long in the Rocky
Mountain district, short in the Pacific district (see figs. 2, 3)--is
paralleled in the _Penthestes atricapillus_ group of chickadees across
the North American continent in about the latitude of the state of
Washington. In the northern Rocky Mountains occurs the race _P. a.
septentrionalis_, with long tail; in the Pacific Coast strip of
Washington and Oregon occurs the race _P. a. occidentalis_, with, among
other characters, relatively short tail. Other parallels are to be found
in the genera _Psaltriparus_, _Thryomanes_ and _Pipilo_.
[Illustration: Fig. 2. Tail of _Penthestes gambeli gambeli_
(no. 27784, Mus. Vert. Zool.; [Male], full-grown juv.;
Sierra Ancha, Gila Co., Arizona; June 26, 1917). × 1.00.]
[Illustration: Fig. 3. Tail of _Penthestes gambeli abbreviatus_
(no. 24059, Mus. Vert. Zool.; [Male], full-grown juv.; Yolla
Bolly Mt., Tehama Co., California; August 5, 1913). × 1.00.]
It is improbable that the fact of subspecific differentiation in
_Penthestes gambeli_ has been altogether overlooked previous to the
present time. A premonition of the geographic variation now formally
pointed out is to be found in the early writings of Baird (1858, p.
394), who remarked that "a specimen, apparently of this species [_Parus
montanus_], from Medicine Bow creek [Wyoming], (5643,) though marked
female, is larger than those from California, as shown by the
measurements." Also, some of the differences in proportions in the
subspecies now recognized are shown in the table of measurements given
by Ridgway (1904, p. 409).
For a detailed description of the Mountain Chickadee as a _species_,
the reader is referred to Ridgway (1904, p. 408).
=Penthestes gambeli gambeli= (Ridgway)
Rocky Mountain Chickadee
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