Of all the changes that have taken place in organic life, none perhaps
are more remarkable than the transformations which the Cephalopod
molluscs have undergone during the various geological eras. In the more
ancient Palæozoic seas flourished the Orthoceratites, or straight-
chambered shells, resembling a nautilus uncoiled. In the Carboniferous
ages the Goniatites acquired their highest development. These shells
were spirally wound, having the lobes of the chambers free from lateral
denticulations or crenatures, so as to form continuous and uninterrupted
outlines.
Both Orthoceratites and Goniatites disappear in the Triassic times, and
are replaced by hosts of Ammonites, which successively flourished in
more than 600 species, and are characterised by an external siphon and
chambers of complicated, often foliated, pattern. This foliated
structure gives a remarkable character to the intersection of the
chamber partitions with the shell, and must have added greatly to the
strength of the shell, which was always delicate and often very
beautiful. The Ammonites, which made their first appearance towards the
end of the Triassic period, abounded in the Oolitic and Cretaceous
periods, and were replaced by new forms before the Tertiary beds were
deposited. Among these we find the _Ancyloceras gigas_, which may be
regarded as an Ammonite partially unrolled, and the _Turrilites
tuberculatus_, which has the form and peculiar symmetry of a univalve
shell.
[Illustration: TURRILITES TUBERCULATUS.]
[Illustration: RESTORED BELEMNITE.]
In several of the older rocks, especially the Lias and Oolite,
Belemnites are frequently met with. These singular dart- or arrow-shaped
fossils were supposed by the ancients to be the thunderbolts of Jove,
but are now known to be the petrified internal bones of a race of
voracious cuttle-fishes, whose importance in the Oolitic or Cretaceous
Seas may be judged of by the frequency of their remains and the 120
species that have been hitherto discovered.
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