The Systematics of the Frogs of the Hyla Rubra Group in Middle America — John Shaqi
The Systematics of the Frogs of the Hyla Rubra Group in Middle AmericaLeón, Juan R.
Science
The Systematics of the Frogs of the Hyla Rubra Group in Middle America
León, Juan R.
Frogs -- Central America; Hyla rubra
_Hyla boulengeri:_ Günther, Biologia Centrali-Americana, Reptilia
and Batrachia, p. 267, June 1901. Noble, Bull. Amer. Mus. Nat.
Hist, 38:339, June 1918. Taylor, Univ. Kansas Sci. Bull., 35:856,
July 1, 1952.
_Diagnosis._--Size large (Male to 49 mm., Female to 53 mm.); skull as
long as wide; frontoparietal fontanelle present; snout subacuminate;
canthus not pronounced; choanae large; tongue cordiform, slightly
longer than broad; interorbital triangle tubercular; skin on dorsum
tuberculate; tarsal fold reduced or absent; thighs, shanks, and tarsi
boldly barred with dark brown and pale yellow-green in life.
_Description._--Head flattened, longer than wide; snout projecting
beyond lower lip; loreal region oblique; canthus not pronounced; length
of eye less than interorbital distance; tympanum large, about 70 per
cent of diameter of eye; interorbital triangle distinct; arms short;
fingers lacking web; palmar tubercle tripartite; subarticular tubercles
distinct; long tubercle on base of first finger; discs truncate; legs
long; tarsal fold reduced or absent; inner metatarsal tubercle rounded,
larger than outer, both elevated; subarticular tubercles distinct; one
phalanx free of web on second, third, and fifth toes, three free on
fourth toe (Fig. 1A and B); skin tuberculate on dorsum, less so on
flanks; skin of belly granular, that on chest and throat weakly
granular; tongue cordiform, longer than wide, free and notched behind;
vocal slits large, lateral to tongue.
[Illustration: Fig. 1. A and B.--Hand and foot of _Hyla boulengeri_
(KU 102173), × 3. C and D.--Hand and foot of _Hyla s. staufferi_× (KU
57790), × 6]
In life, dorsum tan or brown with dark spots on snout, head, and
scapular region; interorbital triangle and blotch posteriorly on dorsum
dark brown; flanks pale green; groin pale green or orange, mottled with
dark brown; thighs tan or brown above with dark transverse bars on
anterior and posterior surfaces; spaces between bars green or orange;
inner surfaces of shanks and outer surfaces of tarsi brown and orange;
foot brown above; forelimbs brown and pale green above, weakly barred;
belly creamy white with scattered brown spots; vocal sac creamy white
flecked with brown; lower jaw brown with white spots on lips (Pl. 1A).
In preservative, head and dorsum dark brown with triangular spot
between eyes; dark spots on head and scapular region and dark brown
blotch posteriorly on dorsum; flanks creamy white with brown spots;
groin creamy white mottled with dark brown; thighs brown above with
dark brown transverse bars on anterior and posterior surfaces; inner
surfaces of shanks and outer surfaces of tarsi barred with pale brown;
dorsal surface of foot mottled brown and creamy white; ventral surface
of foot and toes pale brown; forelimbs faintly barred with pale brown;
belly white with a few pale brown spots; vocal sac flecked with pale
brown; lower jaw marked with small white spots on lips.
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