From these quasi-vertebrates developed the fishes—first (after the scaleless, limbless lampreys) the sharks with spiny scales and cartilaginous skeleton, and after these the lung fishes and the bony fishes, with flat, horny scales and skeletons of bone. From the beginning of the Devonian age, when fishes first came into prominence, till the rise of the great reptiles in the Triassic time, fishes were the dominant life of the sea. In the fishes first appeared jaws, a sympathetic nervous system, red blood, backbone, and the characteristic two pairs of limbs of vertebrates. The lung fishes (Dipneusta), a small order of strange salamander-like creatures which live ingeniously on the borderland between the liquid and the land, may be looked upon as physiological, if not morphological, links between the fishes and the frogs. They combine the characters of both fishes and frogs, and zoologists have been tempted to make a separate class of them, and place them between the two classes to which they are related. They are like fishes in having scales, fins, permanent gills, and a fish-like shape and skeleton. They resemble frogs in having lungs, nostrils, an incipiently three-chambered heart, a pulmonary circulation, and frog-like skin glands. There are three genera with several species. One genus (Neoceratodus) is found in two or three small rivers of Queensland, Australia; another (Protopterus) lives in the Gambia and other rivers of Africa; and the third (Lepidosiren) inhabits the swamps of the Amazon region. They all breathe ordinarily by means of gills, like true fishes, but have the habit of coming frequently to the surface and inhaling air. The air-bladder acts as an incipient lung in supplementing respiration by gills. They all live in regions where a dry season regularly converts the watercourses into beds of sand and mud. During the season of drought these strange animals build for themselves a cocoon or nest of mud and leaves. This cocoon is lined with mucus, and provided with a lid through which air is admitted. Here they lie in this capsule throughout the hot southern summer, from August to December, breathing air by means of their lungs and living upon the stored-up fat of their tails, until the return of the wet season, when they again live in the rivers and breathe water in true piscatorial fashion. These capsules have often been carried to Europe, and opened 3,000 miles from their place of construction without harming the life within.