increase in size at each moult, so that the male ultimately becomes
spotted with black. With Carriers, the wattle, both on the beak and
round the eyes, and with Barbs that round the eyes, goes on
increasing with age. This augmentation of character with advancing
age, and more especially the difference between the males and
females in the above-mentioned several respects, are remarkable
facts, for there is no sensible difference at any age between the
two sexes in the aboriginal rock-pigeon; and not often any strongly
marked difference throughout the family of the Columbidæ.[35]
_Osteological Characters._
In the skeletons of the various breeds there is much variability; and
though certain differences occur frequently, and others rarely, in
certain breeds, yet none can be said to be absolutely characteristic of
any breed. Considering that strongly-marked domestic races have been
formed chiefly by man’s selection, we ought not to expect to find great
and constant differences in the skeleton; for fanciers neither see, nor
do they care for, modifications of structure in the internal framework.
Nor ought we to expect changes in the skeletons from changed habits of
life; as every facility is given to the most distinct breeds to follow
the same habits, and the much modified races are never allowed to
wander abroad and procure their own food in various ways. Moreover, I
find, on comparing the skeletons of _Columba livia, oenas, palumbus,_
and _turtur_, which are ranked by all systematists in two or three
distinct though allied genera, that the differences are extremely
slight, certainly less than between the skeletons of some of the most
distinct domestic breeds. How far the skeleton of the wild rock-pigeon
is constant I have had no means of judging, as I have examined only
two.
Illustration: Fig. 24—Skulls of Pigeons, viewed laterally.
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