It has already been remarked that dogs differ in the degree to which their
feet are webbed. In dogs of the Newfoundland breed, which are eminently
aquatic in their habits, the skin, according to Isidore Geoffroy,[78]
extends to the third phalanges, whilst in ordinary dogs it extends only to
the second. In two Newfoundland dogs which I examined, when the toes were
stretched apart and viewed on the under side, the skin extended in a nearly
straight line between the outer margins of the balls of the toes; whereas,
in two terriers of distinct sub-breeds, the skin viewed in the same manner
was deeply scooped out. In Canada there is a dog which is peculiar to the
country and common there, and this has "half-webbed feet and is fond of the
water."[79] English otter-hounds are said to have webbed feet: a friend
examined for me the feet of two, in comparison {40} with the feet of some
harriers and bloodhounds; he found the skin variable in extent in all, but
more developed in the otter than in the other hounds.[80] As aquatic
animals which belong to quite different orders have webbed feet, there can
be no doubt that this structure would be serviceable to dogs that frequent
the water. We may confidently infer that no man ever selected his
water-dogs by the extent to which the skin was developed between their
toes; but what he does, is to preserve and breed from those individuals
which hunt best in the water, or best retrieve wounded game, and thus he
unconsciously selects dogs with feet slightly better webbed. Man thus
closely imitates Natural Selection. We have an excellent illustration of
this same process in North America, where, according to Sir J.
Richardson,[81] all the wolves, foxes, and aboriginal domestic dogs have
their feet broader than in the corresponding species of the Old World, and
"well calculated for running on the snow." Now, in these Arctic regions,
the life or death of every animal will often depend on its success in
hunting over the snow when softened; and this will in part depend on the
feet being broad; yet they must not be so broad as to interfere with the
activity of the animal when the ground is sticky, or with its power of
burrowing holes, or with other habits of life.
Public-domain text, read in full here on John Shaqi.
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