he tried in vain during several years to get seed from these flowers. I
likewise made many vain attempts, but sometimes succeeded in fertilising them
with pollen from a normal flower of another variety; and conversely I several
times fertilised ordinary flowers with peloric pollen. Only once I succeeded
in raising a plant from a peloric flower fertilised by pollen from a peloric
flower borne by another variety; but the plant, it may be added, presented
nothing particular in its structure. Hence we may conclude that no general
rule can be laid down; but any great deviation from the normal structure, even
when the reproductive organs themselves are not seriously affected, certainly
often leads to sexual impotence.
DOUBLE FLOWERS.
When the stamens are converted into petals, the plant becomes on the male side
sterile; when both stamens and pistils are thus changed, the plant becomes
completely barren. Symmetrical flowers having numerous stamens and petals are
the most liable to become double, as perhaps follows from all multiple organs
being the most subject to variability. But flowers furnished with only a few
stamens, and others which are asymmetrical in structure, sometimes become
double, as we see with the double gorse or Ulex, and Antirrhinum. The
Compositae bear what are called double flowers by the abnormal development of
the corolla of their central florets. Doubleness is sometimes connected with
prolification (18/96. 'Gardener's Chronicle' 1866 page 681.), or the continued
growth of the axis of the flower. Doubleness is strongly inherited. No one has
produced, as Lindley remarks (18/97. 'Theory of Horticulture' page 333.),
double flowers by promoting the perfect health of the plant. On the contrary,
unnatural conditions of life favour their production. There is some reason to
believe that seeds kept during many years, and seeds believed to be
imperfectly fertilised, yield double flowers more freely than fresh and
perfectly fertilised seed. (18/98. Mr. Fairweather 'Transact. Hort. Soc.'
volume 3 page 406: Bosse quoted by Bronn 'Geschichte der Natur' b. 2 s. 77. On
the effects of the removal of the anthers see Mr. Leitner in Silliman's 'North
American Journ. of Science' volume 23 page 47; and Verlot 'Des Varietes' 1865
page 84.) Long-continued cultivation in rich soil seems to be the commonest
exciting cause. A double narcissus and a double Anthemis nobilis, transplanted
into very poor soil, has been observed to become single (18/99. Lindley's
'Theory of Horticulture' page 3?3.); and I have seen a completely double white
primrose rendered permanently single by being divided and transplanted whilst
in full flower. It has been observed by Professor E. Morren that doubleness of
the flowers and variegation of the leaves are antagonistic states; but so many
exceptions to the rule have lately been recorded (18/100. 'Gardener's
Chronicle' 1865 page 626; 1866 pages 290, 730; and Verlot 'Des Varietes' page
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