75.), that, though general, it cannot be looked at as invariable. Variegation
seems generally to result from a feeble or atrophied condition of the plant,
and a large proportion of the seedlings raised from parents, if both are
variegated, usually perish at an early age; hence we may perhaps infer that
doubleness, which is the antagonistic state, commonly arises from a plethoric
condition. On the other hand, extremely poor soil sometimes, though rarely,
appears to cause doubleness: I formerly described (18/101. 'Gardener's
Chronicle' 1843 page 628. In this article I suggested the theory above given
on the doubleness of flowers. This view is adopted by Carriere 'Production et
Fix. des Varietes' 1865 page 67.) some completely double, bud-like, flowers
produced in large numbers by stunted wild plants of Gentiana amarella growing
on a poor chalky bank. I have also noticed a distinct tendency to doubleness
in the flowers of a Ranunculus, Horse-chestnut, and Bladder-nut (Ranunculus
repens, Aesculus pavia, and Staphylea), growing under very unfavourable
conditions. Professor Lehmann (18/102. Quoted by Gartner 'Bastarderzeugung' s.
567.) found several wild plants growing near a hot spring with double flowers.
With respect to the cause of doubleness, which arises, as we see, under widely
different circumstances, I shall presently attempt to show that the most
probable view is that unnatural conditions first give a tendency to sterility,
and that then, on the principle of compensation, as the reproductive organs do
not perform their proper functions, they either become developed into petals,
or additional petals are formed. This view has lately been supported by Mr.
Laxton (18/103. 'Gardener's Chronicle' 1866 page 901.) who advances the case
of some common peas, which, after long-continued heavy rain, flowered a second
time, and produced double flowers.
SEEDLESS FRUIT.
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