prostrate yew (_Taxus baccata_) was found in a hedge in Shropshire; it
was a male, but one branch bore female flowers, and produced berries;
these, {19} being sown, produced seventeen trees, all of which had
exactly the same peculiar habit with the parent-tree.[47]
These facts, it might have been thought, would have been sufficient to
render it probable that a pendulous habit would in all cases be
strictly inherited. But let us look to the other side. Mr. MacNab[48]
sowed seeds of the weeping beech (_Fagus sylvanica_), but succeeded in
raising only common beeches. Mr. Rivers, at my request, raised a number
of seedlings from three distinct varieties of weeping elm; and at least
one of the parent-trees was so situated that it could not have been
crossed by any other elm; but none of the young trees, now about a foot
or two in height, show the least signs of weeping. Mr. Rivers formerly
sowed above twenty thousand seeds of the weeping ash (_Fraxinus
excelsior_), and not a single seedling was in the least degree
pendulous: in Germany, M. Borchmeyer raised a thousand seedlings, with
the same result. Nevertheless, Mr. Anderson, of the Chelsea Botanic
Garden, by sowing seed from a weeping ash, which was found before the
year 1780, in Cambridgeshire, raised several pendulous trees.[49]
Professor Henslow also informs me that some seedlings from a female
weeping ash in the Botanic Garden at Cambridge were at first a little
pendulous, but afterwards became quite upright: it is probable that
this latter tree, which transmits to a certain extent its pendulous
habit, was derived by a bud from the same original Cambridgeshire
stock; whilst other weeping ashes may have had a distinct origin. But
the crowning case, communicated to me by Mr. Rivers, which shows how
capricious is the inheritance of a pendulous habit, is that a variety
of another species of ash (_F. lentiscifolia_) which was formerly
pendulous, "now about twenty years old has long lost this habit, every
shoot being remarkably erect; but seedlings formerly raised from it
were perfectly prostrate, the stems not rising more than two inches
above the ground." Thus the weeping variety of the common ash, which
has been extensively propagated by buds during a long period, did not,
with Mr. Rivers, transmit its character to one seedling out of above
twenty thousand; whereas the weeping variety of a second species of
ash, which could not, whilst grown in the same garden, retain its own
weeping character, transmitted to its seedlings the pendulous habit in
excess!
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