The law of reversion is equally powerful with hybrids, when they are
sufficiently fertile to breed together, or when they are repeatedly crossed
with either pure parent-form, as with mongrels. It is not necessary to give
instances, for in the case of plants almost every one who has worked on
this subject from the time of Koelreuter to the present day has insisted on
this tendency. Gaertner has recorded some good instances; but no one has
given more striking cases than Naudin.[83] The tendency differs in degree
or strength in different groups, and partly depends, as we shall presently
see, on the fact of the parent-plants having been long cultivated. Although
the tendency to reversion is extremely general with nearly all mongrels and
hybrids, it cannot be considered as invariably characteristic of them;
there is, also, reason to believe that it may be mastered by long-continued
selection; but these subjects will more properly be discussed in a future
chapter on Crossing. From what we see of the power and scope of reversion,
both in pure races and when varieties or species are crossed, we may infer
that characters of almost every kind are capable of reappearance after
having been lost for a great length of time. But it does not follow from
this that in each particular case certain characters will reappear: for
instance, this will not occur when a race is crossed with another endowed
with prepotency of transmission. In some few cases the power of reversion
wholly fails, without our being able to assign any cause for the failure:
thus it has been stated that in a French family in which 85 out of above
600 members, during six generations, had been subject to night-blindness,
"there has not been a single example of this affection in the children of
parents who were themselves free from it."[84]
* * * * *
_Reversion through Bud-propagation--Partial Reversion, by segments in the
same flower or fruit, or in different parts of the {37} body in the same
individual animal._--In the eleventh chapter, many cases of reversion by
buds, independently of seminal generation, were given--as when a leaf-bud
on a variegated, curled, or laciniated variety suddenly reassumes its
proper character; or as when a Provence-rose appears on a moss-rose, or a
peach on a nectarine-tree. In some of these cases only half the flower or
fruit, or a smaller segment, or mere stripes, reassumed their former
character; and here we have with buds reversion by segments. Vilmorin[85]
has also recorded several cases with plants derived from seed, of flowers
reverting by stripes or blotches to their primitive colours: he states that
in all such cases a white or pale-coloured variety must first be formed,
and, when this is propagated for a length of time by seed, striped
seedlings occasionally make their appearance; and these can afterwards by
care be multiplied by seed.
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