The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
In =imperfect joints=, such as the intervertebral joints of mammals,
the two contiguous surfaces are united by a mass of fibrous tissue
which allows only a limited amount of motion.
In =perfect joints= the contiguous articular surfaces are covered with
cartilage, and between them lies a synovial membrane which secretes a
viscid lubricating fluid.
The amount of motion possible varies according to the nature of the
articular surfaces; these include--
_a._ =ball and socket joints=, like the hip and shoulder, in which the
end of one bone works in a cup provided by another, and movements can
take place in a variety of planes.
_b._ =hinge joints=, like the elbow and knee, in which as in
ball-and-socket joints one bone works in a cup provided by another,
but movements can take place in one plane only.
THE ENDOSKELETON.
The endoskeleton is divisible into =axial= and =appendicular= parts;
and the =axial= skeleton into--
1. the spinal column,
2. the skull {_a._ the cranium,
{_b._ the jaws and visceral skeleton,
3. the ribs and sternum[7].
I. THE AXIAL SKELETON.
1. THE SPINAL COLUMN.
The spinal column in the simplest cases consists of an unsegmented
rod, the notochord, surrounded by the =skeletogenous layer=, a sheath
of mesoblastic origin, which also envelops the nerve cord. Several
intermediate stages connect this simple spinal column with the
vertebral column characteristic of higher vertebrates. A typical
vertebral column may be said to consist of (1) a series of
cartilaginous or bony blocks, the vertebral =centra=, which arise in
the sheath surrounding the notochord. They cause the notochord to
become constricted and to atrophy to a varying extent, though a
remnant of it persists, either permanently or for a long period,
within each centrum or between successive centra. (2) From the dorsal
surface of each centrum arise a pair of processes which grow round the
spinal cord and unite above it, forming a =dorsal= or =neural arch=.
(3) A similar pair of processes arising from the ventral surface of
the centrum form the =ventral= or =haemal arch=. To the ventral arch
the ribs strictly belong, and it tends to surround the ventral
blood-vessels and the body cavity with the alimentary canal and other
viscera.
A =neural spine= or spinous process commonly projects upwards from the
dorsal surface of the neural arch, and a pair of =transverse
processes= project outwards from its sides. When, as is commonly the
case, the two halves of the haemal arch do not meet, the ventral
surface of the centrum often bears a downwardly-projecting
=hypapophysis=.
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