The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
(_b_) At the _posterior end_ of the cranium the most prominent feature
is the large, almost circular =foramen magnum=, through which the
spinal cord and brain communicate; this in young birds is seen to be
bounded by four distinct bones, dorsally by the =supra-occipital=,
ventrally by the =basi-occipital=, and laterally by the
=exoccipitals=.
The =basi-occipital= forms the main part of a prominent convex knob,
the =occipital condyle=, with which the atlas articulates. The
occipital condyle is slightly notched above, and the ventral surface
of the cranium is deeply pitted just in front of it; the exoccipitals
also contribute slightly to its formation. Slightly in front of and
ventral to the foramen magnum is a small foramen through which the
hypoglossal nerve leaves the cranial cavity.
The =supra-occipital= is separated from the parietal by a suture line
along which run a pair of prominent ridges, the =lambdoidal crests=
(fig. 60, B, 30). There are often a pair of prominent vacuities in the
supra-occipital dorsal to the foramen magnum. The =epi-otics= and
=opisthotics= become completely fused with the bones of the occipital
segment at a very early stage.
(_c_) The _ventral surface_ of the cranium is wide behind, where it is
formed by a broad transverse membrane bone, the _basitemporal_ (fig.
60, A, 23), the sides of which are fused with the auditory capsules.
Slightly in front of and an eighth of an inch external to the
hypoglossal foramen the cranial wall is pierced by a pair of foramina
through which the tenth or pneumogastric nerves leave (fig. 60, A, X).
At the sides of the basitemporal are a pair of depressions, the
=tympanic recesses=, in each of which are three holes. Straight lines
joining these holes would form an isosceles triangle with its apex
directed forwards. Of the two holes at the base of the triangle, the
one nearer the middle line and leading into the cranial cavity, is for
the exit of the ninth or glossopharyngeal nerve (fig. 60, A, IX), it
lies just in front of the pneumogastric foramen. The more external
leads into the tympanic cavity, while the more anterior at the apex of
the triangle is the =posterior opening of the carotid canal= (fig. 60,
A, 25), which traverses the base of the cranium, and during life
lodges the carotid artery.
The anterior end of the basitemporal is pierced near the middle line
by a pair of holes, the =anterior openings of the Eustachian canals=;
while just in front of these and a little further removed from the
middle line are the anterior openings of the =carotid canals=.
Bristles passed in through the posterior openings of the carotid
canals will emerge here (fig. 60, A, 27). In front of the basitemporal
the base of the cranium is formed by the =rostrum= (fig. 60, A, 31),
or thickened basal portion of the interorbital septum; this bears two
prominent surfaces with which the pterygoids articulate. In some kinds
of duck these surfaces are borne by well-marked basi-pterygoid
processes.
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