The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
(_d_) _The side of the cranium._ At the base of the posterior end is
seen the deep =tympanic cavity=. The dorsal part of this is divided by
a vertical partition into two halves; of these the more anterior is
the larger, and forms a deep funnel-shaped cavity, the =posterior
opening of the Eustachian canal= (fig. 60, B, 26). A bristle passed
into this opening emerges through the anterior opening of the
Eustachian canal. The more posterior of the two is the =fenestral
recess= (fig. 60, B, 28), and is in its turn divided by a slender
horizontal bar into a dorsal hole, the =fenestra ovalis=, and a
ventral hole, the =fenestra rotunda=. During life the fenestra ovalis
lodges the proximal end of the =columellar= chain. Lying at the outer
side and slightly dorsal to the tympanic cavity is a deep depression,
the =lateral tympanic recess=, and immediately in front of this is the
articular surface for the quadrate. The tympanic cavity is bounded
below by the basitemporal, posteriorly by the exoccipital, and above
by the _squamosal_, a membrane bone, which roofs over a good deal of
the side of the cranium, and bears ventrally a prominent surface with
which the quadrate articulates. Just in front of this is a large round
hole, the =trigeminal foramen= (fig. 60, B, V), behind which the
squamosal is drawn out into a short process.
In front of the squamosal there is a prominent forwardly-projecting
=postfrontal process= (fig. 60, 8), which ossifies from a different
centre from that forming the squamosal, but in the adult is completely
fused with it.
The =orbit= forms a large more or less hemispherical cavity which
lodges the eyeball. It is separated from its fellow of the opposite
side by an imperfect partition, the =interorbital septum=. In the
young skull it is seen to be bounded above by the frontal, with which
the _lachrymal_ (fig. 60, 7) is fused anteriorly, forming a large
backwardly-projecting process; while behind it is bounded by
the =alisphenoid=. The interorbital septum is formed by the
ossification and coalescence of the =mesethmoid= in front, with the
=orbitosphenoid= behind, and the =rostrum= below. The boundary of the
orbit below is very imperfect, the zygomatic arch being incomplete.
The interorbital septum is pierced by the very prominent =optic
foramen= (fig. 60, B, 2), just behind which are the two much smaller
foramina for the exit of the oculomotor and pathetic (fig. 60, B, IV)
nerves, the more anterior being that for the oculomotor.
Above and slightly in front of the optic foramen is a median opening,
the =olfactory foramen.= This leads into the cranial cavity behind,
and in front is continued forwards as a groove between the
interorbital septum and the frontal.
(2) THE FACIAL PART OF THE SKULL.
This includes the olfactory capsule and associated bones, and the
upper jaw.
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