The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
The bones associated with the olfactory capsules are the _nasals_ and
_vomer_. The _nasals_ (figs. 59 and 60, 5) lie on the dorsal surface
immediately in front of the cranium, and are separated from one
another by the nasal processes of the premaxillae. Each is completely
fused in the adult with the corresponding maxillae and premaxillae, the
three bones together forming the boundary of the =anterior nares.= The
_vomer_ (fig. 60, 19) is unpaired and forms a small median vertical
plate lying ventral to the anterior continuation of the interorbital
septum.
The bones of the upper jaw consist on each side of two slender arcades
which in front converge and are attached to the large beak, while
behind they diverge but are united by the =quadrate=.
The =inner arcade= is formed by the pterygoid and palatine. The
_pterygoid_ (fig. 60, 21) is a short flattened bone, which articulates
behind with the quadrate, and on its inner side with a large flattened
surface borne by the rostrum, in front it meets the palatine, or
sometimes ends freely with a long antero-dorsally directed point.
The _palatine_ (fig. 60, 20) is a slender irregular bone flattened
dorso-ventrally at its anterior end where it articulates with the
beak, and laterally behind. It gives off at its posterior end a
process, which is sometimes united with the vomer, sometimes projects
forwards, and meets its fellow dorsal to the vomer. In the large space
between it and the vomer is the opening of the =posterior nares=.
The _premaxillae_ (figs. 59 and 60, 2) are very large, and form nearly
a third of the big shovel-shaped beak. They constitute the inner, and
part of the front boundary of the anterior nares, and send back a pair
of _nasal processes_ which partially separate the nasals from one
another.
The =outer arcade= forms the slender =suborbital bar=, and consists
mainly of two rod-like bones, which in the adult are completely fused
together. The posterior of these is the _quadratojugal_ (figs. 59 and
60, 11) which articulates with the quadrate, the anterior is the small
and slender _jugal_ or _malar_ (figs. 59 and 60, 10). The extreme
anterior part of the bar is formed by the _maxillae_. The main part of
the maxillae however lies anterior to the suborbital bar, and extends
forwards along the side of the premaxillae forming all the lateral part
of the beak (figs. 59 and 60, 1); it also sends inwards a plate, the
=maxillo-palatine= (fig. 60, A, 29), which completely fuses with its
fellow in the middle line, and forms the posterior boundary of the
anterior palatine foramen. The term =desmognathous= describes the
condition of the skull in which the maxillo-palatines fuse with one
another in the middle line in this way.
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