The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
In higher vertebrates ossification sets up in the cartilage and gives
rise on each side to a dorsal bone, the =scapula=, and frequently to
an anterior ventral bone, the =precoracoid=, and a posterior ventral
bone, the =coracoid=. The precoracoid is often not ossified, and upon
it is developed the clavicle which more or less replaces it. In some
forms a =T= shaped _interclavicle_ occurs, in others =epicoracoids= are
found in front of the coracoids. In all vertebrata above fish, except
the great majority of mammals, the coracoids are large and articulate
with the sternum. But in mammals the coracoids are nearly always quite
vestigial, and the pectoral girdle is attached to the axial skeleton
by the clavicle or sometimes by muscles and ligaments only.
The =Pelvic girdle=[12] like the pectoral consists primitively of a
simple rod or hoop of cartilage, which in vertebrata above fishes is
divided into dorsal and ventral portions, by a cavity, the
=acetabulum=, with which the posterior limb articulates. In the pelvic
girdle as in the pectoral one dorsal, and (commonly) two ventral
ossifications take place. The dorsal bone is the =ilium= and
corresponds to the scapula. The posterior ventral bone is the
=ischium= corresponding to the coracoid. The anterior ventral bone is
the =pubis= and is generally compared to the precoracoid, but in some
cases a fourth pelvic element, the =acetabular= or =cotyloid= bone is
found, and this may correspond to the precoracoid.
The pelvic girdle differs from the pectoral in the fact that the
dorsal bones--the ilia--are nearly always firmly united to transverse
processes of the sacral vertebrae, by means of rudimentary ribs. The
pubes and ischia generally meet in ventral symphyses.
2. THE LIMBS.
It will be most convenient to defer a discussion of the limbs of
fishes to chap. VIII.
All vertebrates above fishes have the limbs divisible into three main
segments:--
=Anterior or Fore limb.= =Posterior or Hind limb.=
Proximal segment. upper arm or _brachium_. thigh.
Middle segment. fore-arm or _antibrachium_. shin or _crus_.
Distal segment. _manus_. _pes_.
The proximal segments each contain one bone, the =humerus= in the case
of the upper arm, and the =femur= in the case of the thigh. The middle
segments each contain two bones, the =radius= and =ulna= in the case
of the fore-arm, and the =tibia= and =fibula= in the case of the shin.
* * * * *
The manus and pes are further subdivided into
(_a_) two or three proximal rows of bones forming the wrist or
=carpus= in the case of the manus, and the ankle or =tarsus= in the
case of the pes.
(_b_) a middle row called respectively the =metacarpus= and
=metatarsus=.
(_c_) a number of distal bones called the =phalanges= which form the
skeleton of the fingers and toes, or =digits=.
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