The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
In the Sperm whale, _Physeter_, the lower jaw bears a series of twenty
to twenty-five stout conical recurved teeth, while in the upper jaw
the teeth are vestigial and remain imbedded in the gum. An extinct
form, _Physodon_, from the Pliocene of Europe and Patagonia is allied
to the Sperm whale, but has teeth in both jaws. In the Grampus _Orca_,
the teeth number about 12/12, and are very large and strong. In some
forms the teeth are very much reduced in number; thus in _Mesoplodon_
the dentition consists simply of a pair of conical teeth borne in the
mandible. In the Narwhal _Monodon_ the dentition is practically
reduced to a single pair of teeth, which lie horizontally in the
maxillae, and in the female normally remain permanently in the alveoli.
In the male the right tooth remains rudimentary, while the left is
developed into an enormous cylindrical tusk marked by a spiral groove.
Occasionally both teeth develop into tusks, and there is reason for
thinking that two-tusked individuals are generally or always female.
In the extinct _Squalodon_ the dentition is decidedly heterodont, and
the molars have two roots. The dental formula is
_i_ 3/3 _c_ 1/1 _pm_ 4/4 _m_ 7/7, total 60.
It is probable that the homodont condition of modern Odontoceti is not
primitive, but due to retrogressive evolution.
UNGULATA.
Just as in the Cetacea a piscivorous dentition is most typically
developed, so the Ungulata are, as a group, the most characteristic
representatives of a _herbivorous_ dentition in its various forms.
UNGULATA VERA.
_ARTIODACTYLA_. As regards the living forms, the Artiodactyla can be
readily divided into two groups, namely those with bunodont and those
with selenodont teeth. It has, however, been shown that selenodont
teeth always pass through an embryonic bunodont stage[157]. The
bunodont type is best seen in Pigs and Hippopotami and such extinct
forms as _Hyotherium_. In _Hippopotamus_ the dental formula is _i_
(2-3)/(1-3) _c_ 1/1 _pm_ 4/4 _m_ 3/3.
The incisors and canines of _Hippopotamus_ are very large and grow
continuously. The genus _Sus_, which affords a good instance of an
_omnivorous_ type of dentition, has the regular unmodified Mammalian
dental formula _i_ 3/3 _c_ 1/1 _pm_ 4/4 _m_ 3/3, total 44. The
canines, specially in the male, are large and have persistent pulps,
and the upper canines do not have the usual downward direction but
pass outwards and upwards. In the Wart Hog, _Phacochaerus_, they are
enormously large, but a still more extraordinary development of teeth
is found in _Babirussa_. In the male _Babirussa_ the canines, which
are without enamel, are long, curved and grow continuously. Those of
the upper jaw never enter the mouth, but pierce the skin of the face
and curve backwards over the forehead. The dental formula of
_Babirussa_ is _i_ 2/3 _c_ 1/1 _pm_ 2/2 _m_ 3/3, total 34.
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