The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
In both the living genera the dentition is monophyodont. In _Manatus_
the dentition is _i_ 2/2 _pm_ and _m_ 11/11. The incisors are
vestigial, and disappear before maturity. The grinding teeth have
square enamelled crowns marked by transverse tuberculated ridges. They
are not all present in the jaw at the same time. In _Halicore_ the
upper jaw bears a pair of straight tusklike incisors; in the male
these have persistent pulps and project out of the mouth; in the
female they soon cease to grow and are never cut. They are separated
by a long diastema from the grinding teeth which have tuberculated
crowns and are 5/5 or 6/6 in number, but are not all in place at once.
Several other pairs of slender teeth occur in the young animal, but
are absorbed or fall out before maturity. In _Rhytina_ teeth are
altogether absent.
CETACEA.
_ARCHAEOCETI._ _Zeuglodon_ has the following dentition, _i_ 3/3 _c_
1/1 _pm_ and _m_ 5/5, total 36. The incisors and canines are simple
and conical; the cheek teeth are compressed and have serrated cutting
edges like those in some seals.
In the _MYSTACOCETI_, or whalebone whales, calcified tooth germs
probably belonging to the milk dentition are present in the embryo,
but they are never functional, and are altogether absent in the adult.
The anterior of these germs are simple, the posterior ones are
originally complex, but subsequently split up into simple teeth like
those of the anterior part of the jaw. Hence according to Kükenthal,
who described these structures, the Cetacean dentition was originally
heterodont.
In the living _ODONTOCETI_ the dentition is homodont and monophyodont.
In some cases traces occur of a replacing dentition which never comes
to maturity, and renders it probable that the functional teeth of the
Odontoceti are really homologous with the milk teeth of other mammals.
Some of the dolphins afford the apparently simplest type of mammalian
dentition known. The teeth are all simple, conical, slightly recurved
structures, with simple tapering roots and without enamel. The
dentition is typically _piscivorous_, being adapted for seizing active
slippery animals such as fish. The prey is then swallowed entire
without mastication. Sometimes the teeth are excessively numerous,
reaching two hundred or more (fifty to sixty on each side of each jaw)
in _Pontoporia_. This multiplication of teeth is regarded by Kükenthal
as due to the division into three parts of numbers of trilobed teeth
similar to those of some seals.
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