The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
(3) the skeleton of the =jaws and hyoid apparatus=. The skull is much
flattened and expanded, though not so much as in the frog.
(1) THE CRANIUM PROPER.
The =cranium proper= or =brain case= is an unsegmented tube which
remains partly cartilaginous, and is partly converted into cartilage
bone, partly sheathed by membrane bone. The roof and floor of the
cartilaginous cranium are, as is the case also in the frog, pierced
by holes or fontanelles, and these are so large that the main part of
the roof and floor comes to be formed by membrane bone.
Two pairs of large ossifications take place in the cranial walls. Of
these the more posterior on each side represents the =exoccipital= and
all three =periotic= bones. It bears a small convex patch of cartilage
for articulation with the atlas, and with its fellow forms the
boundary of the foramen magnum.
Two foramina pierce the exoccipital just in front of the occipital
condyle and transmit respectively the glossopharyngeal and
pneumogastric (fig. 21, X) nerves. Lying laterally to these nerve
openings is seen a patch of cartilage, the =stapes=, which is
homologous with the stapes or proximal element of the columellar chain
in the frog. Further forward in front of the stapes is the small
opening for the exit of the facial nerve, and seen in a lateral view
close to the orbitosphenoid, that for the trigeminal (fig. 21, C, 5).
In front of these large bones the lateral parts of the cranial walls
remain cartilaginous for a short distance, and then there follow two
elongated bones, the =orbitosphenoids= (fig. 21, B and C, 11), pierced
by the foramina for the exit of the optic nerves. These bones partly
correspond to the sphenethmoid of the frog.
The _membrane bones_ connected with the cranium are the _parietals_,
_frontals_ and _prefronto-lachrymals_ on the dorsal surface, and the
_parasphenoid_ on the ventral surface.
The _parietals_ (fig. 21, A and C, 6) roof over the posterior part of
the great dorsal fontanelle and overlap the exoccipito-periotics. They
meet one another along a sinuous suture in the middle line, as do also
the _frontals_ which overlap them in front. The _frontals_ and
_parietals_ both extend for a short distance down the sides of the
cranium and meet the orbitosphenoids. The _prefronto-lachrymals_ (fig.
21, A and C, 7) connect the frontals with the maxillae.
[Illustration FIG. 21. A DORSAL, B VENTRAL, AND C LATERAL VIEWS OF THE
SKULL OF A NEWT (_Molge cristata_) × 2-1/2 (after PARKER).
The cartilage is dotted, the cartilage bones are marked with dots and
dashes, the membrane bones are left white.
Public-domain text, read in full here on John Shaqi.
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