The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
To the two groups of unarticulated and articulated sprouts in the
kingdom of the tissue-plants correspond, in many respects, the two
sections of the tissue-animals, the unarticulated and the articulated.
The two stems of the articulates and vertebrates rise above all the
other metazoa by the perfection of their organism and the variety
of their functions. In the articulates the metamerism is chiefly
external--an articulation of the body wall. In the vertebrates it
mainly affects the internal organs, the skeleton, and the muscular
system. The vertebration (articulation) of the vertebrates is not
outwardly visible like that of the articulates. In both stems the
articulation is similar in the lower and upper forms, as we find in
the annelids and myriapods, the acrania and cyclostoma. On the other
hand, the higher the organization the greater is the unlikeness of
the members or articulated parts, as in the arachnida and insects,
the amphibia and amniotes. The same antithesis is found in the lower
and higher crustacea. This metamerism of the higher metazoa is of a
motor character, having been acquired through the manner of movement
of the lengthened body; but we find in some groups of the lower, and
usually unarticulated, metazoa a propagative metamerism, determined
by budding at the end; such is the strobilation of the chain-worms
and the scyphostoma polyps. The individual metamera (parts) that are
released from the end of the chain in these cases immediately show
their individuality. This is also the case with many of the annelids,
in which every member that is separated has the power to reproduce the
whole chain of metamera.
The third and highest stage of individuality to which the multicellular
organism attains is the stock or colony (_cormus_). It is usually
formed by a permanent association of histonals that are produced
by cleavage (imperfect segmentation or budding) from one histonal
individual. The great majority of the metaphyta form complex plants in
this sense. But among the metazoa we find this form of individuality
only in the lower (and generally stationary) stages of development.
Here also there is a striking parallelism of development between the
two chief groups of the histona. At the lower stages of stock-formation
there is equality of the social histonals. But in the higher grades
they become unequally developed in the division of labor; and the
greater the differences between them become, the greater is the
centralization of the whole stock (as in the case of the siphonophora).
We may therefore distinguish two principal forms of stocks--the
homonomous and heteronomous, the one without, and the other with,
division of labor among the histonals.
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