The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
The humerus presents a large convex oval head, on each side of which is
a tuberosity for the implantation of the supra- and sub-scapular
muscles: these tuberosities do not rise above the articular convexity,
so as to restrict the movements of the shoulder-joint, as in the Horse
and Ruminants, but exhibit a structure and disposition conformable to
those which characterize the proximal extremity of the humerus in other
mammalia which enjoy rotatory movements of the upper or fore-limb. The
tuberosities are, however, relatively more developed, and give greater
breadth to the proximal end of the humerus in the Scelidothere than in
the Megathere. The distal end of the humerus, although mutilated,
clearly indicates that it had the same characteristic breadth of the
external and internal condyles, as in the Megatherium. In fig. 1. Pl.
XXV. which gives a front view of the left humerus, the broad internal
condyle, with its extremity broken off, is seen projecting to the left
hand; both in this figure and in fig. 2. in which the internal side of
the humerus is turned towards the observer, the wide groove, with its
two osseous boundaries, is shewn, which plainly indicates that the left
condyle was perforated for the direct passage of the artery or median
nerve, or of both, to the fore-arm. The groove for the musculo-spiral
nerve on the outer side of the humerus is over-arched at its upper part
by a strong obtuse process; which is comparatively less developed in the
Megatherium. The trochlear or inferior articular surface of the humerus
presents, as in the Megatherium, two well-marked convexities, with an
intervening concavity: this indication of the rotatory power of the
fore-leg is confirmed by the form of the head of the radius.
In Pl. XXV. fig. 4. a view is given of this articular surface: it
presents the form of a subcircular gentle concavity, which plays upon
the outer convexity of the humeral articular surface: immediately below
the upper concavity the radius presents a lateral smooth convex surface,
which rotates upon a small concavity on the ulna, analogous to the
‘lesser semilunar,’ in human anatomy, in which the mechanism for
rotation, so far as the upper joint of the radius is concerned, is not
more elaborately wrought out than in the present extinct edentate
quadruped. The radius expands as it proceeds to the elbow-joint, where
it attains a breadth indicative of the great power and size of the
unguiculate paw, of which it may be called the stem, and to the
movements of which it served as the pivot.
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