The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
All the bones of the fore-limb just described—the scapula, the humerus,
and the radius,—indicate by the bold features and projections of the
muscular ridges and tubercles the prodigious force which was
concentrated upon the actions of the fore-paw, and the ulna, in its
broad and high olecranon (of which a side view is given in fig. 2. Pl.
XXV.) gives corresponding evidence. The great semilunar concavity is
traversed by a sub-median smooth ridge, which plays upon the interspace
of the two humeral convexities. The body of the bone is subcompressed,
straight, and diminishes in size as it approaches the carpal joint: the
immediate articulating surfaces are wanting in both the radius and ulna,
the epiphyseal distal extremities having become detached from their
respective diaphyses.
Of the terminal segment of the locomotive extremities, the only evidence
among the remains of the skeleton of the Scelidothere is the ungueal
phalanx figured at Pl. XXVII. 3, 4, and 5; but as it is uncertain
whether it belongs to the fore or hind-foot, it will be described after
the other bones of the extremities have been noticed.
Of these bones the femur is the most remarkable, both for its great
proportional size, and its extreme breadth, as compared with its length
or thickness: but in all these circumstances the affinity of the
Scelidothere with the Megathere is prominently brought into view. There
is no other known quadruped with which the Scelidothere so closely
corresponds in this respect. In proceeding, however, to compare together
the thigh-bones of these two extinct quadrupeds, several differences
present themselves, which are worthy of notice: of these the first is
the presence in the Scelidothere of a depression for a ‘ligamentum
teres’ on the back part of the head of the femur, near its junction with
the neck of the bone: this is shewn in the posterior view of the femur
given in Pl. XX. The head itself forms a pretty regular hemisphere: the
great trochanter does not rise so high as in the Megatherium, but,
relatively, it emulates it in breadth: the small trochanter is
proportionally more developed: the external contour of the shaft of the
femur is straighter in the Scelidothere than in the Megathere, and the
shaft itself is less bowed forwards at that part. The articular condyles
occupy a relatively smaller space upon the distal extremity of the femur
in the Scelidothere, and they differ more strikingly from those of the
Megathere, in being continued one into the other: the rotular surface,
for example, which is shewn in fig. 5. Pl. XXV. is formed by both
condyles, while in the Megatherium it is a continuation exclusively of
the external articular surface.
The patella, which works upon the above-mentioned surface, is a thick
strong ovate bone, with the smaller end downwards: rough and convex
externally, smooth on the internal surface, which is concave in the
vertical and convex in the transverse directions.
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