The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
In the Armadillo the astragalus, in consequence of the greater
production of its anterior part, presents more of an angular than a
quadrate figure; and the scaphoid articular surface, being
proportionally carried forwards, is altogether separated from the
anterior calcaneal surface. The posterior and inner calcaneal surface
resembles that in the Scelidothere, but is less inclined upwards; and is
continuous with the posterior part of the tibial articular surface.
Thus the astragalus in the structure of its two most important
articulations, viz. that which receives the superincumbent weight from
the leg, and that which transmits it to the heel, presents a closer
correspondence in the Scelidothere with that of the Dasypus, than with
that of the Megathere or Mylodon.
The ungueal phalanx of the Scelidothere before alluded to, is
represented of the natural size in Pl. XXVII. The side view, fig. 3.
shows the position of the articular surface on the proximal end, sloping
obliquely towards the under surface, and overtopped by an obtuse
protuberance, calculated to impede any upward retraction of the claw:
the present joint, in fact, illustrates in every particular the argument
by which Cuvier established the true affinities of the allied extinct
genus Megalonyx.[58]
The present phalanx is, however, less compressed, and less incurved than
those of the Megalonyx, which have been hitherto described; but it more
resembles in these proportions one of the smaller, and presumed hinder,
ungueal phalanges of the Megatherium. The upper and lateral parts of the
bone are rounded, and it gradually tapers to the apex, which is broken
off. The osseous sheath for the claw is developed only at the under part
of the bone: it presents the form of a thick flat plate of bone, with
the margin very regularly and obliquely bevelled off, and having a
vertical process of bone attached lengthwise to the middle of its under
surface. This process must have served for the insertion of a very
powerful flexor tendon. The figures of this bone preclude the necessity
of any further verbal description.
M. Lund lays most stress upon the argument founded on the inward
inflection of the sole of the foot in the Megalonyx, and appeals with
greatest confidence to this structure in support of his hypothesis of
the scansorial habits of that extinct Edental.[59]
It is quite true that the Quadrumana derive advantage from this position
of the foot in climbing trees, and that it is carried to excess in the
Sloths, which can only apply the outer edge of the foot to the ground.
But we may ask, was the inversion of the sole of the foot actually
carried to such an extent in the _Megalonyx_? And, admitting its
existence in an inferior degree, is it then conclusive as to the
scansorial habits of that species?
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