The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
The cranium, which is the subject of the present description, was found
in the Sarandis, a small stream entering the Rio Negro, and about 120
miles to the N. W. of Monte Video: it had been originally embedded in a
whitish argillaceous earth, and was discovered lying in the bed of the
rivulet, after a sudden flood had washed down part of the bank.
The zoological characters deducible from this cranium, forbid its
association, generically, with any known Mammiferous animal, and it must
therefore be referred to an extinct genus, which I propose to call
_Toxodon_,[9] from the curved or arched form of the teeth, as will
afterwards be described. The specific name, in the absence of other
means of knowing the peculiarities of the animal than those afforded by
the skull, may be most conveniently taken from the district (La Plata),
in which its remains were first discovered.
The dimensions of the cranium of the _Toxodon Platensis_ amply attest
that the animal to which it belonged was of a magnitude attained by few
terrestrial quadrupeds, and only to be compared, in this respect, with
the larger Pachyderms, or the extinct Megatherium. The length of the
skull (of which a base view of the natural size is given in Plate I.) is
two feet four inches: the extreme breadth one foot four inches. The
other requisite admeasurements are given in the table at the conclusion
of this description.
The general form of the skull, as seen from above, is pyriform; but
viewed sideways, and without the lower jaw, it is semi-ovate; it is
depressed, elongate, of considerable breadth, including the span of the
zygomatic arches, but becoming rather suddenly contracted anterior to
them, the facial part thence growing narrower to near the muzzle, which
again slightly expands.
Among the first peculiarities which strike the observer, is the aspect
of the plane of the occipital foramen, and of the occipital or posterior
region of the cranium, the latter of which inclines from below upwards
and forwards at an angle of 50° with the basal line of the skull. This
slope of the back part of the skull is one of the characteristics of the
Dinotherium; it is common to all the Cetacea, and is met with in a
slighter degree in many Rodentia, and in the great Ant-eater and some
others of the Edentate order. The corresponding aspect of the _foramen
magnum_ presents nearly the opposite extreme to man in the occipital
scale, proposed by Daubenton to determine the diversities of the form of
the cranium, as a gage of the intelligence of different animals[10]; and
the indication of the limited capacity of the Toxodon, thus afforded, is
strengthened by the very small proportion, which the bony walls of the
cerebral cavity bear to the zygomatic and maxillary parts of the skull,
and to the size of the vertebral column, as indicated by the condyloid
processes, and foramen magnum.
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