The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
In the Toxodon the position of the incisors, in the same transverse
line, might lead to the inference, that they were opposed by a
corresponding number in the lower jaw; but the numerous examples of
inequality, in the number of incisors, in the upper and lower jaws of
existing mammalia, forbid any conclusion on this point.[12] The sockets
of the small mesial incisors of the Toxodon (_s s_, Pl. III.) gradually
diminish in size, as they penetrate the intermaxillary bones, and we
may, therefore, infer that the pulp was gradually absorbed in the
progress of their development; and that, like ordinary incisors, their
growth was of limited duration, and their lodgment in the jaw effected
by a single conical fang.
I may observe, that the formation of a fang is the necessary consequence
of the gradual absorption of the matrix or pulp of a tooth; for the pulp
continues, as it diminishes in size, to deposit ivory upon the inner
surface of the cavity of the tooth from which it is receding, and the
tooth or fang thus likewise progressively diminishes in size. The
formation of the socket proceeds uninterruptedly, and the bone
encroaching upon the space left by the tooth, closely surrounds the
wasting fang, and affords it a firm support; and thus an inference may
be drawn from the form of the socket alone, as to whether the tooth it
contained had or had not one or more conical fangs, and consequently
whether its growth was temporary or uninterrupted.
Applying this reasoning to the molar teeth of the Toxodon, we infer that
their growth, like those of most of the Phytiphagous Rodents, of the
Megatherium and Armadillo, was perpetual, because their sockets are
continued of uniform size from the open to the closed extremity; and the
molar tooth which is preserved proves the accuracy of the deduction,
inasmuch as its base is excavated by a large conical cavity for the
lodgment of the pulp, the continued activity of which was the
compensation here designed to meet the effects of attrition on the
opposite or grinding surface of the tooth.
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