The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
The _zygomatic_ process of the temporal bone projects boldly outwards at
its commencement, where it is of great strength, and three-sided; the
glenoid cavity extends transversely across the base or inferior surface
of this part; the lateral surfaces converge to form the ridge or upper
boundary of the zygoma. The depth of the glenoid cavity is increased by
a transverse production of bone both before and behind it: the posterior
process (_g_, Pl. II.) descends the lowest, and affords the requisite
defence against backward dislocation of the lower jaw; the pressure of
the condyle against this process is denoted by a well defined,
transversely-ovate, flattened and smooth surface, as if the bone had
been planed down at that part: the anterior transverse boundary is
convex and smooth, and probably formed part of the articulation for the
lower jaw. The lower facet of the zygoma anterior to the glenoid cavity
gradually contracts in breadth, as it advances forward, and at the
distance of three inches from the articular cavity the zygoma changes
from a prismatic to a laminar form. It is at this point that the
zygomatic suture commences, at the lower margin of the arch; whence it
extends directly forwards for more than half its length, and then bends
upwards at a right angle. The zygomatic suture has a similar course in
the Capybara, and Hippopotamus.
The remainder of the zygoma is formed externally by the _malar_ bone (G
Pl. II.), which in its position is intermediate to the Rodent and
Pachydermatous structures. It is not suspended in the middle of the
zygomatic arch, as in the former order; neither does it extend into the
region of the face so far anterior to the orbit as in the Tapir or
Hippopotamus. The exterior line of the malo-maxillary suture defines the
orbit anteriorly; but from this line the maxillary bone extends
backwards, along the inner side of the malar portion of the zygoma,
until it almost reaches the temporo-malar suture; thus abutting by an
oblique surface against nearly the whole internal facet of the malar
bone, and materially contributing to the general strength of the
zygomatic arch. The malar bone is of considerable vertical extent, and
presents a rugged and thickened inferior margin for the attachment of
the masseter. The upper margin of the malar bone is smoothly rounded,
and presents a regular semicircular excavation, forming the lower
boundary of the orbit. The relative magnitude of the zygomata to the
entire cranium far exceeds in the Toxodon that which exists in the
Hippopotamus or any other known Pachyderm. This arises from the great
vertical development of the malar bone behind the orbit, and the
vertical expansion of the temporal portion of the arch. The oblique
position of the zygoma, descending as it advances forwards, is deserving
of attention, as the Toxodon, in deviating from the Pachyderms in these
respects, makes an evident approach to the herbivorous Cetaceans, as the
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