The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
The most perfect of these fragments is figured in Pl. V. figures 1 and
4; figure 2 shows the form of the teeth in transverse section, and the
disposition of the enamel upon the grinding surface of the molars on the
right side, as restored from a comparison of the fractured teeth in the
two rami. From the remains of the symphysis shown at fig. 4, it will be
seen that the jaw was remarkably compressed, or narrow from side to
side; while the rami (fig. 1.) were of considerable depth, in order to
give lodgment to the matrices and bases of grinders enjoying
uninterrupted growth.
The pulps of the six incisors in this lower jaw are arranged in a pretty
regular semicircle, whose convexity is downwards; the teeth themselves
are directed forwards, and curved upwards, like the inferior incisors of
the Rodentia. The form and degree of the curvature are shown in the
almost perfect incisor (Pl. V. fig. 5) which corresponds with the left
inferior incisor of the lower jaw, and was found in the same stratum,
but belonged to another individual.
These incisors are nearly equal in size: they are all hollow at their
base, and the indurated mineral substance impacted in their basal
cavities well exhibits the form of the vascular pulps which formerly
occupied them. Sufficient of the tooth itself remains in four of the
sockets to show that these incisors, like the nearly perfect one (fig.
5), had only a partial investment of enamel; but though in this respect,
as well as in their curvature and perpetual growth, they resemble the
dentes scalprarii of the Rodentia, they differ in having a prismatic
figure, like the inferior incisors of the Sumatran Rhinoceros, or the
tusks of the Boar. Two of the sides, viz., those forming the anterior
convex and mesial surfaces of the incisor have a coating of enamel,
about half a line in thickness, which terminates at the angles between
these and the posterior or concave surface. In plate V. fig. 4, the
enamel of the broken incisors is represented by short lines, showing the
direction of its crystalline fibres; the white space immediately within
the enamel shows the thickness of the ivory at the base of the tooth,
the included gray substance represents a section of the formative matrix
or pulp of the tooth, which was of the usual conical form: the inferior
broken end of the incisor (fig. 5,) appears to have been distant about
one-third from the apex of the pulp.
From the relative position of the bases or roots of these incisors, we
may infer that they diverged from each other as they advanced forwards,
in order to bring their broadest cutting surfaces into line. That they
were opposed to teeth of a corresponding structure in the upper jaw is
proved by the oblique chisel-like cutting surface of the more perfect
incisor: and it is not without interest to find that the presence of
_dentes scalprarii_ at the anterior part of the mouth has not been
necessarily limited to Mammalia of small size.
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