The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
three-fourths of the body of the vertebra: they are not continued into
the anterior and inferior transverse processes, as in the Vicugna, but
are separated therefrom by a narrow and shallow groove. The articular,
or oblique processes, closely resemble those of the Auchenia in form,
and in the direction of the articular surfaces; those of the anterior
processes looking inwards and a little upwards; those of the posterior,
outwards and a little downwards.
In the Macrauchenia a small longitudinal process (_c_, fig. 2, Pl. VII.)
is given off immediately below the base of the anterior oblique process;
this structure is not observable in any of the cervical vertebræ of the
Giraffe or Camelidæ.
In the form of the articulating surfaces of the bodies of the vertebræ
the Macrauchenia deviates from the Giraffe and Camel, but resembles the
Aucheniæ. In the Giraffe and Camel the anterior articulating surface is
convex and almost hemispheric, the posterior surface is proportionally
concave, so that the cervical vertebræ are articulated by ball and
socket joints; yet not, as in most Reptiles, with intervening synovial
cavities, but by the concentric ligamentous intervertebral substance
characteristic of the Mammiferous class. In the Llama and Vicugna, the
degree of convexity and concavity in the articular surface of the bodies
of the cervical vertebræ is much less than in the Camels; and in
consequence they carry their necks more stiffly and more in a straight
line. In Macrauchenia the anterior articulating surface (fig. 2, Pl.
VII.) presents a still slighter convexity than in the Llama (fig. 4, Pl.
VII.), and the posterior surface (fig. 1, Pl. VII.) presents a
correspondingly shallower concavity. The form of the extremities of the
body of the vertebræ, especially of the posterior, is sub-hexagonal, the
breadth being to the depth as eight to five. The sides and under part of
the vertebræ are slightly concave; on the inferior surface there are two
ridges, continued forwards from the posterior margin of the vertebra,
each situated about an inch distant from the middle line; they converge
as they pass forwards, and are gradually lost in the level of the
vertebra; their greatest elevation does not exceed half an inch. In the
Aucheniæ there is a longitudinal protuberance in the mesial line,
instead of the two ridges. The two long cervical vertebræ of the
Macrauchenia are also characterized by the maintenance of an almost
uniform diameter of the body, both in its vertical and transverse
extent; the cervical vertebræ of the Vicugna come nearest to them in
this respect; those of the Camel deviate further in the large excavation
at the under part of the body.
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