Tropical nature, and other essaysWallace, Alfred Russel
History
Tropical nature, and other essays
Wallace, Alfred Russel
Ecology -- Tropics; Natural history -- Tropics; Tropics
We may now follow the two sexes to the remoter island, at a period
when the male had acquired his permanent style of colouring, but
was not quite so large as he subsequently became; while the change
of the female bird had not been half completed. In this small and
comparatively barren island (a mere rock, as it is described by
some authors) there would be no such constant abundance of food,
and therefore no possibility of a large permanent population; while
the climate would not differ materially from that of the larger
island. Variation would therefore be checked, or might be stopped
altogether; and we find the facts exactly correspond to this view.
The male, which had already acquired his colour, remains almost
undistinguishable from his immediate ancestral form; but he is a
little smaller, indicating either that the full size of that form
had not been acquired at the period of migration, or that a slight
diminution of size has since occurred, owing to a deficiency of
food. The female shows also a slight diminution of size, but in
other respects is almost exactly intermediate between the Chilian
and Juan Fernandez females. The colour beneath is light ashy,
the breast-spots are intermediate in size and colour, and the
tail-feathers have a large ill-defined white spot on the end of
the inner web which has only to be extended along the whole web
to produce the exact character which has been acquired in Juan
Fernandez. It seems probable, therefore, that the female bird has
remained nearly or quite stationary since its migration, while
its Juan Fernandez relative has gone on steadily changing in the
direction already begun; and the more distant species geographically
thus appears to be more nearly related to its Chilian ancestor.
Coming down to a more recent period, we find that the comparatively
small and dull-coloured Chilian bird has again migrated to Juan
Fernandez; but it at once came into competition with its red
descendant, which had firm possession of the soil, and had probably
undergone slight constitutional changes exactly fitting it to its
insular abode. The new-comer, accordingly, only just manages to
maintain its footing; for we are told by Mr. Reed, of Santiago,
that it is by no means common; whereas, as we have seen, the red
species is excessively abundant. We may further suspect that the
Chilian birds now pass over pretty frequently to Juan Fernandez,
and thus keep up the stock; for it must be remembered that whereas,
at a first migration, both a male and a female are necessary for
colonization, yet, after a colony is formed, any stray bird which
may come over adds to the numbers, and checks permanent variation
by cross-breeding.
Public-domain text, read in full here on John Shaqi.
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