Tropical nature, and other essaysWallace, Alfred Russel
History
Tropical nature, and other essays
Wallace, Alfred Russel
Ecology -- Tropics; Natural history -- Tropics; Tropics
We find, then, that all the chief peculiarities of the three allied
species of humming-birds which inhabit the Juan Fernandez group of
islands, may be fairly traced to the action of those general laws
which Mr. Darwin and others have shown to determine the variations
of animals and the perpetuation of those variations. It is also
instructive to note, that where the variations of colour and size
have been greatest they are accompanied by several lesser variations
in other characters. In the Juan Fernandez bird the bill has become
a little shorter, the tail-feathers somewhat broader, and the
fiery cap on the head somewhat smaller; all these peculiarities
being less developed or absent in the birds inhabiting Mas-afuera.
These coincident changes may be due, either to what Mr. Darwin
has termed correlation of growth, or to the partial reappearance
of ancestral characters under more favourable conditions, or to
the direct action of changes of climate and of food; but they show
us how varied and unaccountable are the changes in specific forms
that may be effected in a comparatively short time, and by means
of very slight changes of locality.
If now we consider the enormously varied conditions presented
by the whole continent of America--the hot, moist, and uniform
forest-plains of the Amazon; the open llanos of the Orinoco; the
dry uplands of Brazil; the sheltered valleys and forest slopes of
the Eastern Andes; the verdant plateaus, the barren paramos, the
countless volcanic cones with their peculiar Alpine vegetation; the
contrasts of the East and West coasts; the isolation of the West
Indian islands, and to a less extent of Central America and Mexico
which we know have been several times separated from South America;
and when we further consider that all these characteristically
distinct areas have been subject to cosmical and local changes, to
elevations and depressions, to diminution and increase of size, to
greater extremes and greater uniformity of temperature, to increase
or decrease of rainfall; and that with these changes there have
been coincident changes of vegetation and of animal life, all
affecting in countless ways the growth and development, the forms
and colours, of these wonderful little birds--if we consider all
these varied and complex influences, we shall be less surprised at
their strange forms, their infinite variety, their wondrous beauty.
For how many ages the causes above enumerated may have acted upon
them we cannot say; but their extreme isolation from all other
birds, no less than the abundance and variety of their generic and
specific forms, clearly point to a very high antiquity.
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