Tropical nature, and other essaysWallace, Alfred Russel
History
Tropical nature, and other essays
Wallace, Alfred Russel
Ecology -- Tropics; Natural history -- Tropics; Tropics
_The Relations and Affinities of Humming-birds._--The question of
the position of this family in the class of birds and its affinities
or resemblances to other groups, is so interesting, and affords
such good opportunities for explaining some of the best-established
principles of classification in natural history in a popular way,
that we propose to discuss it at some length, but without entering
into technical details.
There is in the Eastern hemisphere, especially in tropical Africa
and Asia, a family of small birds called Sun-birds, which are
adorned with brilliant metallic colours, and which, in shape and
general appearance, much resemble humming-birds. They frequent
flowers in the same way, feeding on honey and insects; and all
the older naturalists placed the two families side by side as
undoubtedly allied. In the year 1850, in a general catalogue of
birds, Prince Lucien Bonaparte, a learned ornithologist, placed
the humming-birds next to the swifts, and far removed from the
Nectarinidæ or sun-birds; and this view of their position has gained
ground with increasing knowledge, so that now all the more advanced
ornithologists have adopted it. Before proceeding to point out the
reasons for this change of view, it will be well to discuss a few
of the general principles which guide naturalists in the solution
of such problems.
_How to Determine Doubtful Affinities._--It is now generally
admitted that, for the purpose of determining obscure and doubtful
affinities, we must examine by preference those parts of an
animal which have little or no direct influence on its habits
and general economy. The value of an organ, or of any detail of
structure, for purposes of classification, is generally in inverse
proportion to its adaptability to special uses. And the reason of
this is apparent, when we consider that similarities of food and
habits are often accompanied by similarities of external form or
of special organs, in totally distinct animals. Porpoises, for
example, are modified externally so as to resemble fishes; yet
they are really mammalia. Some marsupials are carnivorous, and are
so like true carnivora that it is only by minute peculiarities of
structure that the skeleton of the one can be distinguished from
that of the other. Many of the hornbills and toucans have the same
general form, and resemble each other in habits, in food, and in
their enormous bills; yet peculiarities in the structure of the
feet, in the form of the breast-bone, in the cranium, and in the
texture and arrangement of the plumage, show that they have no
real affinity, the former approaching the kingfishers, the latter
the cuckoos. Such structural peculiarities as these have no direct
relation to habits; and they are therefore little liable to change,
when from any cause a portion of the group may have been driven to
adopt a new mode of life. Thus all the Old World apes, however much
they may differ in size or habits, and whether we class them as
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