Tropical nature, and other essaysWallace, Alfred Russel
History
Tropical nature, and other essays
Wallace, Alfred Russel
Ecology -- Tropics; Natural history -- Tropics; Tropics
Another interesting fact in connexion with this subject is the
presence of arborescent forms of Compositæ in so many of the remotest
oceanic islands. They occur in the Galapagos, in Juan Fernandez,
in St. Helena, in the Sandwich Islands, and in New Zealand; but
they are not directly related to each other; representatives
of totally different tribes of this extensive order becoming
arborescent in each group of islands. The immense range and almost
universal distribution of the Compositæ is due to the combination
of a great facility of distribution (by their seeds) with a great
attractiveness to insects; and to the capacity of being fertilized
by a variety of species of all orders, and especially by flies and
small beetles. Thus they would be among the earliest of flowering
plants to establish themselves on oceanic islands; but where insects
of all kinds were very scarce, it would be an advantage to gain
increased size and longevity, so that fertilization at an interval
of several years might suffice for the continuance of the species.
The arborescent form would combine with increased longevity the
advantage of increased size in the struggle for existence with
ferns and other early colonists; and these advantages have led to
its being independently produced in so many distant localities,
whose chief feature in common is their remoteness from continents
and the extreme poverty of their insect life.
As the sweet odours of flowers are known to act in combination
with their colours, as an attraction to insects, it might be
anticipated that where colour was deficient scent would be so also.
On applying to my friend Sir Joseph Hooker for information as to
the odoriferous qualities of New-Zealand plants, he informed me,
that the New-Zealand flora is, speaking generally, as strikingly
deficient in sweet odours as it is in conspicuous colours. Whether
this peculiarity occurs in other islands I have not been able to
obtain information; but we may certainly expect to find it where
colour is so strikingly deficient as in the flora of the Galapagos
Islands.
Public-domain text, read in full here on John Shaqi.
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