Two New Pelycosaurs from the Lower Permian of Oklahoma — John Shaqi
Two New Pelycosaurs from the Lower Permian of OklahomaFox, Richard C.
Science
Two New Pelycosaurs from the Lower Permian of Oklahoma
Fox, Richard C.
Paleontology -- Permian; Pelycosauria
_Discussion._--The Nitosauridae are small primitive edaphosaurs with a
moderately elongate face, sharp subisodont teeth, little development of
canines and few specializations. The jaw is of a primitive type and
articulates on a level with the tooth-row. The palatal dentition is
primitive (Romer, 1956:280). The nitosaurids are thought to be related
to the later Caseidae, and the most obvious structural similarities are
found in the postcranial skeleton (Vaughn, 1958:989). Cranial
resemblances between the families are fewer, but nevertheless indicate
that a nitosaurid-caseid relationship exists.
Vaughn (1958) described a small pelycosaur, _Colobomycter pholeter_
(Eothyrididae, Ophiacodontia) that structurally resembles the Caseidae.
This individual also was obtained from the Fort Sill locality. In
Vaughn's opinion the features of _Colobomycter_ indicate a close
relationship between eothyridids and caseids and the possibility that
the caseids may well have been of eothyridid rather than nitosaurid
derivation.
In view of this historical uncertainty of the relationships between the
Nitosauridae, the Eothyrididae and the Caseidae, it is well to consider
how the maxillary fragments described above differ from and resemble
representatives of each of these three families as reported in the
literature.
_Delorhynchus_ resembles _Colobomycter_ in size. The mean
extra-maxillary length of the undamaged teeth of the three fragments is
2.5 mm., equal to that reported by Vaughn (1958:985) for teeth about
midway in the postcanine series of _Colobomycter_. None of the teeth of
_Delorhynchus_ extends beyond the maxillary rim as far as does the
canine of _Colobomycter_ (3.5 mm.).
The teeth in both genera are conical and sharply pointed. The naris in
each is enlarged, and the lacrimal is excluded from the narial margin in
each (by inference in _Delorhynchus_.)
The differences between the maxillae of _Colobomycter_ and
_Delorhynchus_ are most striking in the lack of canines in the latter
and the correlated absence of modifications of the maxillary for support
of canines. Additionally, _Delorhynchus_ bears an infraorbital canal in
contrast to the groove in similar position in _Colobomycter_. The
recurvature of the four teeth present in the fragments of _Delorhynchus_
differs from that in the teeth of _Colobomycter_ in which only the
canine and precanine are recurved. Vaughn implies that anterior and
posterior cutting edges extend the length of the teeth in
_Colobomycter_; these are restricted to the distal third of the teeth in
_Delorhynchus_. The external surfaces of the maxillae of _Delorhynchus_
are pitted and ridged; Vaughn was unable to discern sculpturing of the
corresponding surfaces in _Colobomycter_.
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