As to the corolla, it was long since noticed that prolification was especially liable to occur in double flowers; indeed, Dr. Hill, who published a treatise on this subject, setting forth the method of artificially producing prolified flowers, deemed the doubling to be an almost necessary precursor of prolification;[128] but, though frequently so, it is not invariably the case that the flower so affected is double--_e.g._ _Geum_. If double, the doubling may arise from actual multiplication of the petals, or from the substitution of petals for stamens and pistils, according to the particular plant affected. Occasionally in prolified flowers the parts of the corolla, like those of the calyx, become foliaceous, and in the case of proliferous pears fleshy and succulent. There is in cultivation a kind of _Cheiranthus_? in which there is a constant repetition of the calyx and corolla, conjoined with an entire absence of the stamens and pistils; a short internode separates each flower from the one above it, and thus frequently ten or a dozen of these imperfect flowers may be seen on the end of a flower-stalk, giving an appearance as if they were strung like beads, at regular intervals, on a common stalk. I have seen a similar instance in a less degree in a species of _Helianthemum_. The stamens are subject to various changes in prolified flowers; they assume, for instance, a leaf-like or petal-like condition, or take on them more or less of a carpellary form, or they may be entirely absent; but none of these changes seem to be at all necessarily connected with the proliferous state of the flower. Of more interest is the alteration in the position of these organs which sometimes necessarily accrues from the elongation of the axis and the disjunction of the calyx; thus, in proliferous roses the stamens become strictly hypogynous, instead of remaining perigynous. In _Umbelliferae_ the epigynous condition is changed for the perigynous, &c.