The condition of the pistillary organs in prolified flowers has already been alluded to. Hitherto those instances have been considered in which either the carpels were absent, or the new bud proceeded from between the carpels. There is also an interesting class of cases where the prolification is strictly intra-carpellary; the axis is so slightly prolonged that it does not protrude beyond the carpels, does not separate them in any way, but is wholly enclosed within their cavity. Doubtless, in many cases, this is merely a less perfect development of that change in which the axis protrudes beyond the carpels. This intra-carpellary prolification occurs most frequently in plants having a free central placenta, though it is not confined to them, as it is recorded among _Boragineae_. A remarkable instance of this is described by Mr. H. C. Watson in the first volume of Henfrey's 'Botanical Gazette,' p. 88. In this specimen a raceme of small flowers was included within the enlarged pericarp of a species of _Anchusa_. But the most curious instances of this form of prolification are, no doubt, those which are met with among _Primulaceae_ and other orders with free central placentation. Duchartre, in his memoir on the organogeny of plants with a free central placenta, in the 'Ann. des Sc. Nat.,' 3 ser., 1844, p. 290, among other similar instances, mentions two flowers of _Cortusa Matthioli_, wherein the placenta was ovuliferous at the base; but the upper portion, instead of simply elongating itself into a sterile cone, had produced a little flower with its parts slightly different from those of the normal flowers. M. Alph. de Candolle has likewise described somewhat similar deviations, and one in particular in _Primula Auricula_, where the elongated placenta gave off long and dilated funiculi bearing ovules, while other funiculi were destitute of these bodies, but were much dilated and foliaceous in appearance.[129] In some flowers of _Rhododendron_ I have observed a similar condition of the ovules, which, moreover, in the primary flowers, were attached to the walls of the carpels--parietal placentation.