Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
No thalattosaurs were large animals. If they had the same proportions
between the lengths of head, body, and tail as the mosasaurs, none
exceeded seven feet in length, and they may have been even shorter,
though probably not much. The figure of the skull, as restored by
Professor Merriam, shows many striking aquatic adaptations, in the
elongated, pointed muzzle, in the large external nostrils, situated
far back toward the eyes, and in the well-ossified ring of bones
surrounding the eyeball. There is a parietal opening in the roof of
the skull, as in the modern lizards and tuatera; but it is not known
for certainty whether there were two openings on each side in the
roof of the skull, as in the modern tuatera. While this character may
seem trivial, it is really one of the most important in the reptilian
anatomy in determining the relationship and classification of reptiles.
The teeth are conical and pointed in the front end of the upper and
lower jaws, but farther back they are rounded, rugose, and obtuse, and
could have been used only for crushing hard objects, like mollusks,
crustaceans, etc. (Fig. 82). And not only was there a row of such teeth
on each jaw (only partly seen in the figure), but similar teeth covered
a large part of the palate. And the lower jaws, it is seen, are rather
massive.
The vertebrae were, of course, of the more primitive kind, that is,
with the ends concave, both in front and behind. It would have been
strange indeed were they of any other kind, since reptiles with
ball-and-socket joints to the vertebrae, that is, concave on one end
and convex on the other, as in nearly all living reptiles, did not come
into existence till long after the thalattosaurs had disappeared from
geological history; and it is also a curious fact that such vertebrae
appear to have originated only among animals crawling on land, so that
they would not have been a character acquired by the thalattosaurs
after descending into the water. It will be seen from the figure of
a dorsal vertebra that the rib was attached by a single articular
surface, almost exclusively to the body of the vertebra, quite like
those of all lizards, snakes, and mosasaurs, and unlike those of
other reptiles. This too may seem to be a trivial character to prove
relationships with the lizards, but it is a curious fact that no two
animals having different kinds of ribs are closely related to each
other. Possibly, however, this looser mode of attachment of the ribs
in the thalattosaurs was one of their peculiar adaptations to a water
life, and may not have been derived from their land ancestors.
[Illustration: FIG. 83.—Dorsal vertebra of _Thalattosaurus_. (After
Merriam.)]
[Illustration: FIG. 84.—_Thalattosaurus_: bones of front extremity:
_s_, scapula; _c_, coracoid; _h_, humerus; _r_, radius; _u_, ulna.
(After Merriam.)]
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