Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
The Phytosauria, so far as known, were all reptiles of considerable
size, greatly resembling the crocodiles, and especially the gavials
in form and habit, but differing very greatly in having the external
nostrils situated far back near the eyes; in having no false palate
so characteristic of the Crocodilia; in having a more primitive
shoulder-girdle, consisting of a short coracoid, interclavicle, and
clavicles; and in having the ordinary type of pelvis, that is, with
the pubis entering into the acetabular articulation for the femur.
They were all, like the crocodiles, covered more or less by a bony
armor; there are two openings on each side of the temporal region;
there is no pineal opening; the vertebrae are gently biconcave,
precisely like those of the early or mesosuchian crocodiles; there
is always an opening of considerable size, called the preorbital
foramen, in front of the eyes, as in some crocodiles, many dinosaurs,
and most pterodactyls; there is also an opening through the back
part of the mandibles as in crocodiles; and the double-headed ribs
are attached exclusively to the transverse process of the arch,
precisely as in the crocodiles, dinosaurs, and pterodactyls. From
all these it is evident that the phytosaurs are related most nearly
to the crocodiles and dinosaurs, and are probably an early branch
of the stem from which they, the pterodactyls and the birds, arose,
a branch that persisted only a short time, geologically speaking,
and went entirely out of existence at the close of Triassic times,
leaving no descendants behind. Nevertheless, in this comparatively
brief life-span they developed not a few distinctive forms and became
widely distributed over the earth. Their remains are known from the
Upper Trias of Germany, England, and Scotland, India, South Africa,
and from Massachusetts, North and South Carolina, and many places
in the Rocky Mountains. No true phytosaurs are yet known from South
America, but in all probability they will be discovered there when
the Triassic deposits of that continent have been better explored
for fossils. In the Rocky Mountains, especially, their remains are
widely scattered, they have been found in many localities in Wyoming,
Colorado, Oklahoma, Utah, and New Mexico. Though for the most part
their known remains from these localities are yet fragmentary, not
less than four distinct genera have been described from these regions:
“_Belodon_,” _Angistorhinus_, _Paleorhinus_, and _Episcoposaurus_.
From the Carolinas and Massachusetts a single genus, though described
under numerous names, has been made known, originally called by Emmons
_Rutiodon_ (_Rhytidodon_). And from Europe and India at least as many
more different genera are known. All these genera are, however, so
closely allied that they are placed in the single family Belodontidae.
[Illustration: FIG. 92.—Restoration of _Mystriosuchus_, an Upper
Triassic phytosaur.]
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