Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
[Illustration: FIG. 93.—Skull of _Mystriosuchus_, a phytosaur: _pm_,
premaxilla; _m_, maxilla; _na_, nasal; _f_, frontal; _p_, prefrontal;
_l_, lacrimal; _pf_, postfrontal; _po_, postorbital; _pa_, parietal;
_sg_, squamosal; _qj_, quadratojugal; _pl_, palatine; _t_, transverse;
_in_, internal nares; _en_, external nares; _pt_, pterygoid; _bs_,
basisphenoid; _eo_, exoccipital. (After McGregor.)]
[Illustration: FIG. 94.—Dorsal vertebrae of phytosaur: _az_, anterior
zygapophysis; _pz_, posterior zygapophysis; _d_, _c_, articulations of
rib.]
[Illustration: FIG. 95.—Scapula and coracoid of _Rutiodon
carolinensis_, an American phytosaur. (After McGregor.)]
[Illustration: FIG. 96.—_Belodon_; restoration of head, from above.]
[Illustration: FIG. 97.—_Mystriosuchus_: restoration of head, from
above.]
In _Belodon_ (Fig. 96), the earliest known and most typical genus,
perhaps, the moderately elongated face has a high crest reaching nearly
to its front end, and this type is known both from Europe and from New
Mexico. Others have the face long and slender, even longer and more
slender than in the ancient teleosaur crocodiles or the modern gavials.
In some forms the teeth are cylindrical and slender throughout, and
there may be as many as fifty on each jaw, or two hundred in all; while
in others only the anterior teeth are cylindrical and the posterior
teeth are flattened and serrate along their cutting edges. In the body
not very great differences have been observed. Some are more slender
than others, and there are minor differences in the shapes and sizes
and numbers of the bony scutes along the back and on the throat.
But they are all alike in their essential characters, a very long
beak with numerous teeth; the foremost ones on the expanded, more
or less spoon-shaped front extremity, are more or less, sometimes
greatly, elongated. The jaws may be likened to a long and slender
pair of tongs with nipping teeth at the front end. The strong, long,
and flattened tail is sufficient evidence that the phytosaurs were
excellent swimmers, but, aside from that and the posterior location of
the external nostrils, directly over the internal, few other aquatic
adaptations are observed in the skeleton. There are no sclerotic bony
plates about the eyes, or at least none have so far been discovered,
although among the numerous known specimens they would confidently be
expected were they really present in the skeleton; and the presence of
bony armor negatives markedly aquatic habits.
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