Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
The following scheme differs only in minor details from the more
conservative of the generally accepted views, and those differences
are, for the most part, the writer’s own opinions, to be taken for
what they are worth. It may be said decisively that no classification
of the reptiles into major groups, into super-families or subclasses
that has so far been proposed is worthy of acceptance; there is no
such subclass as the Diapsida or Synapsida, for instance. And we have
very much more to learn about the early reptiles before any general
classification of the reptiles can be securely founded. It is very
probable that the primary radiation of the reptiles into the various
lines of descent, into its main branches, occurred much earlier than
we have been disposed to believe; that before the close of Paleozoic
time, perhaps before the close of the Carboniferous, all the great
groups of reptiles had gone off from the main stem, and that since then
only smaller and smaller branches have appeared. There have been no new
orders of reptiles in all probability since Triassic times, and perhaps
none since Permian.
Taxonomists are often disposed to cut the Gordian knots of
relationships by raising the ranks of the animals they study to
independent positions. More than thirty independent orders of reptiles
have been proposed by different students, and quite as many of mammals
and of birds; possibly after more forms have been discovered there will
be as many proposed for the amphibians. Sometimes, indeed, it is better
to make such independent groups than to unite lesser ones on doubtful
evidence. But the writer, for one, believes that it is more worthy of
the thoughtful scientific student to seek for relationships than for
differences. It is far easier to destroy than to construct, to make
new genera, families, and orders than to unite those already proposed.
To raise every proposed suborder of reptiles to an order, as has been
proposed by various writers, and the orders to subclasses, only leaves
classification where it was; nothing has been added to taxonomy save a
lot of new names to perplex and annoy the student.
In the following scheme of classification three groups provisionally
called orders are prefixed by an asterisk.
CLASS REPTILIA
Order =COTYLOSAURIA=
Primitive reptiles with notochordal vertebrae, imperforate
temporal region, persistent intercentra; two coracoids;
plate-like pelvis, with all or most of the amphibian skull
elements; short legs and short neck; phalangeal formula
primarily 2, 3, 4, 5, 3(4).
Suborder =Diadectosauria= Permocarboniferous, North America.
=Pantylosauria= Permocarboniferous, North America.
=Labidosauria= Lower Permian, North America.
=Pareiasauria= Upper Permian, Europe, Africa.
=Procolophonia= Triassic, Europe, Africa.
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