Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
While the ancient crocodiles of which we have spoken resembled the
modern ones so closely in form of body and probably in habits, there
were certain others of the old Jurassic seas which departed so widely
both in structure and in habits, from their associates that they are by
some authors given a place wholly by themselves as a distinct group.
This has been called by Professor Fraas the Thalattosuchia, a word
meaning “sea-crocodiles.” They were a very early side-branch from the
great genealogical tree of the Crocodilia, a branch which departed so
widely from their associates in adapting themselves to a peculiar and
aberrant mode of existence that they cannot be considered as typical
crocodiles, although so closely related to them in other respects
that therecannot be the least doubt regarding their ancestry. None of
the crocodiles which we have considered, whether ancient or modern,
can truthfully be called purely aquatic. They never ceased to use
their limbs for locomotion on land, more or less of the time, or for
the support of the body; and many of them have subsisted, in part at
least, on land animals. How easy it may have been for some of them
to become almost wholly emancipated from land habits we may easily
conjecture. The gavials, as we have seen, are more at home in the
water than upon land; their food is chiefly found in the water; but,
so long as their habits restrict them chiefly to fresh-water, or to
the vicinity of the shores, their limbs continue to be used as much
for crawling as for swimming. Were the gavials to be driven out to
sea by the stress of fresh-water conditions or attracted thereto by a
greater abundance of more easily obtainable or better food, so far from
land that they no longer would have much use for their legs for the
support or propulsion of their bodies, in the course of time they would
doubtless change to a more purely aquatic type. And in that change
there would be material modifications of their structure: their limbs
would become better adapted to movements in the water; the skin would
become smoother, and the bony and horny scales would be lost, since
they would be not only useless in the water, but actually detrimental
to the well-being of the animals; and the tail would develop into a
more powerful organ of propulsion, as a means of increasing their speed
in obtaining food or in escaping their enemies, such as the sharks.
In fact, we can only imagine that in the transformation precisely
those modifications would occur which we actually find in these old
sea-crocodiles, the Thalattosuchia. And they are of especial interest
to us here because nowhere do we find a better example of Nature in the
act of transforming a terrestrial or subterrestrial animal into a truly
aquatic one.
[Illustration: FIG. 107.—_Geosaurus_, an Upper Jurassic thalattosuchian
crocodile, drawn from restoration and figures by Fraas.]
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