Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
The group comprises only a few forms, so far as known. All were of
modest size among extinct reptiles, from ten to twenty feet in length,
and all are from the Upper Jurassic deposits of Europe. They did
not exist very long, probably because they found it impossible to
discard old habits and old structures entirely and become absolutely
emancipated from the land; their breeding habits possibly were too
deeply impressed into their structure readily to change, as did those
of other sea-reptiles. Some of their remarkable aquatic adaptations
have long been known, but only within a dozen years has our knowledge
of them become at all complete. Three or four genera have been
described, but only a few forms are well known, of which _Geosaurus_
may be taken as most typical. To this we shall confine our descriptions.
[Illustration: FIG. 108.—_Geosaurus_; skull from side and from above.
(After Fraas.)]
The skull of _Geosaurus_ is rather small in comparison with the length
of the body, smaller proportionally than in any living crocodile, but
not much smaller than that of the teleosaurs. The snout is long and
slender, much like that of the teleosaurs and gavials, but the bones
of the whole upper surface are quite smooth, not roughened and pitted
like those of modern forms. The skull of _Dakosaurus_, another genus
of thalattosuchians, is much less elongate than that of _Geosaurus_,
but has the other characteristics of _Geosaurus_. The eyes are provided
with a stout ring of sclerotic bones, with a pupillary opening of less
than one inch. We have seen that all other strictly aquatic reptiles
have similar eye bones, but no other crocodiles have them. The internal
openings of the nostrils are large and long, but they are not situated
far back, as in the modern crocodiles, not even so far back as in the
early teleosaurs. They had no need of the peculiar breathing apparatus
of the amphibious crocodiles, since all their prey must have been
water-breathing creatures. Their eyes were directed laterally, not
more or less upward, as in their nearest relatives. Nearly all other
crocodiles have an opening through the hind end of the lower jaw, but
the thalattosuchians did not. The teeth were about as numerous as in
the modern gavials, but they projected freely only a short distance
above the gums in life, and they were very slender and sharply pointed,
excellently well adapted for catching smooth and slippery fishes.
Their vertebrae, like those of all other reptiles of their time,
were biconcave. Those of the neck resembled those of the teleosaurs,
save that there were only seven, fewer than is the case with any
other members of the order. In becoming adapted to their peculiar
mode of life these crocodiles lost two vertebrae from the neck. All
modern crocodiles have two ribs attached to the first vertebra; the
thalattosaurs had but one, another evidence of primitive characters.
While the number of vertebrae in the neck was reduced, in the back it
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