Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
was increased to eighteen; all other crocodiles have but fifteen or
sixteen. The trunk was long, another adaptation to water life. There
were two firmly united vertebrae in the sacrum, as in the modern forms.
The reason for the persistence of this terrestrial character we shall
see later.
[Illustration: FIG. 109.—Tail, scapula (_sc_), and coracoid (_c_) of
_Geosaurus_. (After Fraas.)]
The tail was very long and strong, nearly as long as all the remainder
of the body, and relatively much longer than in other crocodiles. It is
of interest to observe that the head, neck, body, and tail had almost
the same relative proportions as those of the great sea-lizards, the
mosasaurs. The terminal bones of the tail are very peculiar, and very
different from the corresponding bones of other crocodiles. While the
spines of the tail bones along the anterior part are only moderately
stout and long, and are directed obliquely backward, near the terminal
part they become suddenly much broader and are directed upward, and, a
little farther along, obliquely forward. The chevron bones on the under
side also here become broader and longer. The end of the tail curves
markedly downward to end in a slender point. It will be remembered that
a similar downward curvature of the end of the tail observed in nearly
all specimens of ichthyosaurs induced in Owen the belief that the
animals had a fleshy terminal fin, a belief which later discoveries of
the carbonized remains of the flesh confirmed. The peculiar structures
observed in various specimens of these sea-crocodiles, even though no
impressions or remains of the fleshy parts have been discovered, is
quite conclusive evidence that these animals also had a broad, fleshy,
terminal fin. No other explanation of the structure is possible.
[Illustration: FIG. 110.—_Geosaurus._ Elongate hind leg, and
paddle-like front leg. (After Fraas.)]
The ribs are not at all stout and are not much curved. They are
directed posteriorly in the known specimens preserved in the matrix
in such a way as to indicate a slender thorax and abdomen, not the
broad body of the modern crocodiles. The abdominal ribs, that is,
those protecting the region on the under side of the body between the
breast bone and the pelvis, are strongly developed in _Geosaurus_. The
sternum, always present in other crocodiles, is wanting in _Geosaurus_.
The shoulder-blades and coracoids, often changed in shape in water
reptiles, are not unlike those of the amphibious crocodiles, but are
smaller and flatter.
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