Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
[Illustration: FIG. 10.—Mandible of _Labidosaurus_, a cotylosaur
reptile: _A_ from within; _B_ from without. The coronoid (_cor_) is a
single bone, but extends far forward. The splenial (_sp_) is also a
single bone, replacing the two of the amphibians. The _prearticular_
(_pa_) is narrower, and the angular (_ang_) appears broadly on the
inner side. The dentary (_d_) is much larger and the surangular (_sa_)
is distinct. The articular (_art_) is small.]
[Illustration: FIG. 11.—Mandible of _Alligator_, a modern, highly
specialized reptile, from within. The coronoid (_cor_) is small and
is situated far back; the splenial (_sp_) does not extend to the
symphysis; the prearticular (_pa_) has disappeared, or has fused with
the angular (_an_) or articular (_art_). The dentary (_d_) has become
the chief bone of the mandible.]
The teeth of reptiles are of much less importance, as a rule, in the
determination of relationships than are the teeth of mammals. Rarely
are their shapes of specific, and often not of generic, importance,
though their number and relative sizes may be. The teeth of mammals,
as a rule, are forty-four or less in number, and they are always
inserted in distinct sockets in the jaw bones. Among reptiles they
are indefinite in number, and may be attached to any of the bones
of the palate and sometimes also to the coronoid of the mandibles.
Furthermore, except in those reptiles related to the immediate
ancestors of the mammals, they are alike or nearly alike in the
jaws, that is, homodont, not distinguishable into incisors, canines,
and molars. They may be inserted in separate sockets (thecodont),
in grooves, or simply be co-ossified to the surface of the bone
(acrodont). And they are usually reproduced indefinitely by new teeth
growing at the side of the base or below them. More usually they are
pointed and curved; sometimes they are flattened, with sharp cutting
edges in front and behind in the more strictly carnivorous reptiles;
in those of herbivorous habits they are more dilated and roughened on
the crown, not pointed; in not a few they are low, broad, and flat
and are used only for crushing the hard shells of invertebrates. With
the very few exceptions among certain dinosaurs, they never have more
than one root for attachment. The evolutional tendency for reptiles,
as for the mammals, is to loose teeth, especially those of the palate.
Among living reptiles it is only the most primitive types, such as
the lizards, snakes, and the tuatera, which have teeth on the palatal
bones, and in none are there teeth on the vomers, as was the rule in
the ancient reptiles. The lizards may have them on pterygoids and
palatines, and the tuatera has them on the palatines only. There may be
as many as eighty on each jaw, above and below, and hundreds of smaller
ones on the palate, or they may be reduced in number to five or six,
or even to a single one; some reptiles, like the turtles and later
pterodactyls, have none.
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