Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
We classify all the known forms of reptiles from the Lower Permian
under three or four orders, the Cotylosauria, Theromorpha or
Pelycosauria, Proganosauria, and possibly the Protorosauria, but the
classification is yet provisional, representing merely the present
stage of our knowledge. The Proganosauria and Protorosauria, including
distinctively aquatic reptiles, will be more fully described in
the following pages. To give even a brief description of the more
terrestrial reptiles of this, the earliest known reptilian fauna, would
be beyond our purpose; the accompanying life restorations by the author
of some of the more typical and better known forms, based upon nearly
perfect skeletons, will suffice.
From the reptiles and amphibians of the Lower Permian of Texas and New
Mexico to the ichthyosaurs of the Middle Triassic of California there
is a complete gap in the records of the land life of North America. We
do not know what became of all the remarkable animals of the Permian.
There are few traces of their descendants elsewhere known, unless it
be in South Africa. From the Middle and Upper Permian of South Africa
and Russia, a marvelous reptilian fauna has been made known in recent
years. More than a hundred species of six or seven groups, and at least
two orders have been described. Of these the Cotylosauria are the
continuation of the American order, but include more specialized forms,
the Pareiasauria and the Procolophonia, all of them, like the more
primitive American forms, characterized by the imperforate temporal
region. The Therapsida, likewise, seem to be the continuation of the
American Theromorpha, so closely allied to them that it is difficult
to draw a distinguishing line between them. On the other hand, these
African reptiles merge through the Theriodontia into the mammals in the
Triassic. They are all terrestrial, crawling reptiles, except a few
which are described on a later page under the Anomodontia.
[Illustration: FIG. 27.—Restoration of _Labidosaurus_, a cotylosaur
reptile from Texas, about three feet long.]
The records of the lower part of the Triassic period are scanty
everywhere in the world, save perhaps in Africa. Before the close of
the period, however, probably every important group of cold-blooded
air-breathing animals had made its appearance in geological history,
if we except the snakes; even the mammals had appeared, and possibly
the birds. The Cotylosauria, Theromorpha, and Therapsida disappeared,
the latter giving birth to the mammals; the nothosaurs and plesiosaurs,
the ichthyosaurs, dinosaurs, crocodiles, phytosaurs, rhynchocephalians,
lizards, and turtles have all left records of their existence in Upper
Triassic rocks; and the pterodactyls had also, in all probability,
begun their career, though none is surely known till the Jurassic.
Public-domain text, read in full here on John Shaqi.
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